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Chapter 34 Phylum Hemichordata- Acorn Worms. Have three body regions: proboscis, collar, and trunk Have both a ventral and dorsal nerve cord anterior portion of dorsal is hollow Have pharyngeal gill slits Have ciliated larvae identical to echinoderms Have a hydrostatic skeleton
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Chapter 34Phylum Hemichordata- Acorn Worms • Have three body regions: proboscis, collar, and trunk • Have both a ventral and dorsal nerve cord anterior portion of dorsal is hollow • Have pharyngeal gill slits • Have ciliated larvae identical to echinoderms • Have a hydrostatic skeleton • Filter-feeders using their pharyngeal slits
Chapter 34 Vertebrate Evolution and Diversity Characteristics of Chordata 1. Notochord 2. Dorsal hollow nerve cord 3. Pharyngeal Slits 4. Muscular Postanal Tail
Notochord: This is a longitudinal, flexible rod of cartilage that is located between the gut and the dorsal nerve cord. The notochord can last into adulthood, but in most cases it is replaced by the vertebral column. The spongy material in between the vertebral bones is all that is left of the notochord.
Dorsal, Hollow Nerve Cord: This is a tube that is formed from a plate of ectoderm that is rolled up into a tube. It is located dorsal to the notochord, it lies above the gut and beneath a single, hollow nerve cord. Due to its hollow structure the dorsal nerve cord is considered very unique. It will develop into the chordate's central nervous system.
Pharyngeal Gill Slits: These structures are openings of the upper digestive tube. During the embryonic stage these slits connect the outside to the pharynx. Early chordates used them to filter feeding. Over time the slits became modified to function in gas exchange.
Muscular Post anal Tail: Most chordates have a tail extending beyond the anus. In many aquatic species the tail is equipped with skeletal and muscular tissue for movement. Anus Tail
Subphylum Urochordata-sea squirtstunicates; cellulose in outer covering
Subphylum Cephalochordata lancelets
Subphylum Cephalochordata lancelets • Most like early chordates (but chordates did not descend from them) • Marine filter feeders • Feeble swimmers • Muscles segments develop from blocks of mesoderm called somites that are arranged around each side of the notochord.
Subphylum Cephalochordata lancelets
Subphylum Cephalochordata lancelets
Fossils of early vertebrates
Subphylum Vertebrata • Notochord develops into a cartilaginous or boney vertebral column that surrounds and protects the spinal cord • Have a cranium or skull that surrounds and protects the brain • Have a advanced closed circulatory system • Cephalization increases dramatically
Embryonic formation of the neural crest
Class Myxini-Jawless fish-Hagfish-All marine No limbs or appendages; Cartilaginous skeleton; no vertebrae
Class Cephalaspidomorphi-lamprey-Jawless fish; Cartilaginous skeleton; No limbs or appendages; notochord has dorsal extensions that partially enclose the nerve cord “vertebrae”
Ostracoderms- armored but no jaws; some were active and had paired fins
Placoderms-first jaws • Skeletal rods called gill arches of the most anterior pharyngeal gill slits became the jaw
Placoderms Placoderms
Chondricthyes sharks and rays • Completely cartilaginous skeleton; • Strong powerful swimmers • Must swim constantly to produce water flow over gills due to no operculum • Lateral line system that detects sound waves • Oviparous • Ovoviviparous • Viviparous • Cloaca
Osteichthyes-bony fish • Operculum-bony covering over gill arches pulls water into mouth and over the gills • Swim bladder- gives buoyancy regulates level in water; evolved from crude lungs • Bony fish evolved in freshwater first and developed crude lungs to supplement gills for gas exchange; then when they returned to salt water, the opening to the lungs closed and they evolved into swim bladders • Two chambered heart