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Predation. One species benefits, the other incurs a cost; also herbivory & parasitism. Important in adaptive evolution: predator and prey adaptations. Orcinus orca calls. Phoca vitulina response (most common prey). Predator (usually silent).
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Predation. One species benefits, the other incurs a cost; also herbivory & parasitism. Important in adaptive evolution: predator and prey adaptations. Orcinus orca calls Phoca vitulina response (most common prey) Predator (usually silent) Changes in number of seals visible at the surface Distance of nearest seal to playback source Familiar non-predator (frequently vocal) Unfamiliar non-predator (frequently vocal) Nature 2002 R. W. Baird pages 1178-1179 Plant defenses against herbivory. Also important in shaping communities. K. Starks
Mutualism.Both species benefit from the interaction. Some are obligatory: both species cannot persist without the other; others are facultative: the association is nonessential. Overlooked as a mechanism in community structure; it may be more significant than either competition or predation. F M Bill H. bihai H. caribaea Flower H. bihai H. bihai H. caribaea Inflorescence St. Lucia H. bihai H. caribaea H. caribaea Inflorescence Dominica Science 2003 Also important in adaptive evolution pages 1180-1181
SOME questions from February 17th 1- When determining human population growth, do the only consider natural death or all causes? 2- In the barnacle example, are you saying that Balanus is better suited to the environment and that if desiccation was not a problem at the higher water level, they would force out the Chthamalus? 3- Assuming two species occupy the same ecological niche, why couldn’t they both co-exist assuming they were equally fit?
Species Interactions Competition, Predation, Mutualism, Commensalism [Coevolution: reciprocal evolutionary change in interacting species, ending in some degree of specialization.] Trophic Structure Dominant and Keystone Species Community Control COMMUNITY STRUCTURE
TROPHIC STRUCTURE Feeding relationships between organisms. Describe species interactions. Food chain- Transfer of food energy all the way until decomposers. Trophic levels- Links in the food chain. Usually four or five. Food web- Branching and interconnected food chains. Fig. 53.10 page 1181
ANTARCTIC FOOD WEB page 1182 Fig. 53.11
FOOD WEB TUESDAY LAKE, MI Fish Zooplankton Phytoplankton Proc. Natl. Acad. Sci. 18 Feb 2003
Productivity in holes on tree trunks Fig. 53.13 What limits the length of a food chain? Energetic hypothesis. Inefficiency of energy transfer along the chain limits the number of levels. Dynamic stability hypothesis. Long food chains are less stable than short food chains. page 1183
For example, sugar maple and impact on shading and soil and thus forest community. treeguide.com Species that control community structure Dominant species. Have the highest abundance or highest biomass (sum weight of all individuals). Exert control over the distribution and abundance of other species. pages 1183-1184
Keystone species. Exert strong control on community structure by their ecological roles (niches). They are not particularly abundant. For example, Pisaster ochraceous (ochre sea star) and Mytilus californianus (mussel). Fig. 53.14 Keystone in wave-exposed sites, not so in wave-protected sites due to low food supply and sand burial. pages 1184-1185
Top-down model of regulation of community structure pages 1185-1186 western Alaska Science 1998
Examples of changes in community structure regulated by predators Bioscience 2005
Bottom-up model of regulation of community structure Mar. Ecol. Prog. Ser. 2005 pages 1185-1186
Disturbance-Data suggest that the norm for most communities IS disturbance and nonequilibrium (constantly changing), rather than stability and equilibrium (species composition relatively constant). Disturbances are events that damage communities, remove species and alter resource availability. Fig. 53.17 pages 1186-1188
Mount St. Helens 1980 1978 ÙSFS Despite the importance of natural disturbances, humans are the greatest agents of disturbance. 1999 1981
Ecological succession- Transitions in species composition over ecological time. Primary succession- Begins in a virtually lifeless area where soil has yet not formed (new volcanic island or rubble left by retreating glaciers). Glacier Bay, Alaska Spruce butting in: 80-200 yrs. Alder, cottonwoods: 30-80 yrs. Spruce and hemlock: 200-300 yrs. Secondary succession- Occurs where an existing community has been cleared by some disturbance that leaves the soil intact.
Secondary succession- Occurs where an existing community has been cleared by some disturbance that leaves the soil intact. Yellowstone One year later