Chapter 49: Sensory and Motor Mechanisms

1. Chapter 49:Sensory and Motor Mechanisms By: Nicole Huffman Period 7

2. Key Terms • Sensations: action potentials that reach the brain via sensory neurons. • Perception: constructions formed in the brain and do not exist outside it such as colors, smells, sounds, and tastes. • Sensory Reception: the detection of a stimulus by sensory cells. • Sensory Receptors: are specialized neurons or epithelial cells that exist singly or in groups with other cell types in sensory organs. • Exteroreceptors: sensory receptors that detect stimuli coming from outside the body. • Interoreceptors: detect stimuli coming form within the body, such as blood pressure and body position.

3. Key Terms Continued… • Sensory Transduction: the conversion of stimulus energy into a change in the membrane potential of a sensory receptor. • Receptor Potential: the change in membrane potential itself. • Amplification: the strengthening of stimulus energy by cells in sensory pathways. • Sensory Adaptation: a decrease in responsiveness during continued stimulation. • Mechanoreceptors: sense physical deformation caused by stimuli such as pressure, touch, stretch, motion, and sound – all forms of mechanical energy.

4. Key Terms Continued… • Muscle spindles: dendrites of sensory neurons that spiral around the middle of small skeletal muscle fibers each containing 2 to 12 of these fibers surrounded by connective tissue, parallel to other muscle fibers. • Chemoreceptors: include both general receptors that transmit information about the total solute concentration of a solution and specific receptors that respond to individual kinds of molecules. • Electromagnetic Receptors: detect various forms of electromagnetic energy, such as visible light, electricity, and magnetism. • Photoreceptors: electromagnetic receptors that detect the radiation known as visible light.

5. Key Terms Continued… • Thermoreceptors: respond to heat or cold, help regulate body temperature by signaling both surface and body core temperature. • Pain Receptors/Nociceptors: a class of naked dendrites in the epidermis. • Lateral Line System: mechanoreceptors that detect low-frequency waves by a mechanism similar to the function of the inner ear. • Compound eyes: consists of several thousand light detectors called ommatidia. • Sclera: a touch, white outer layer of connective tissue.

6. Key Terms Continued… • Choroid: a thin, pigmented inner layer • Conjunctiva: A delicate layer of epithelial cells forms a mucous membrane that covers the outer surface of the sclera and helps keep the eye moist. • Cornea: At the front of the eye and lets light into the eye and acts as a fixed lens. • Iris: The anterior choroid that gives the eye its color. • Pupil: the hold in the center of the iris.

7. Key Terms Continued… • Retina: the innermost layer of the eyeball and contains the photoreceptors. • Ciliary Body: produces the clear watery aqueous humor that fills the anterior cavity. • Vitreous Humor: fills the posterior cavity and constitutes most of the volume of the eye. • Fovea: the center of the visual field. • Retinal: a light-absorbing molecule.

8. Key Terms Continued… • Opsin: a membrane protein. • Rhodopsin: a visual pigment in rods. • Photopsins: the three visual pigments of cones. • Ganglion cells: synapse with bipolar cells and transmit action potentials to the brain via axons in the optic nerve. • Horizontal cells and Amacrine cells: help integrate the information before it is sent to the brain.

9. Key terms continued… • Lateral Inhibition: sharpens edges and enhances contrast in an image. • Hydrostatic Skeleton: a skeleton consisting of fluid held under pressure in a closed body compartment. • Skeletal Muscle: muscle attached to the bones and is responsible for their movement. • Accommmodation: the focusing of light in the retina.

10. Weakmuscle stretch Strongmuscle stretch Muscle Dendrites Receptor potential –50 –50 –70 –70 Stretchreceptor Membranepotential (mV) Action potentials 0 0 Axon –70 –70 4 5 1 2 6 7 0 0 1 2 3 4 5 6 7 3 Time (sec) Time (sec) receptor. A stronger stretch produces a larger receptor potential and higher frequency of action potentials. (a) Crayfish stretch receptors have dendrites embedded in abdominal muscles. When the abdomen bends, muscles and dendrites stretch, producing a receptor potential in the stretch receptor. The receptor potential triggers action potentials in the axon of the stretch Figure 49.2a Sensory receptors

11. No fluidmovement Fluid moving inone direction Fluid moving in other direction “Hairs” ofhair cell Neuro-trans-mitter at synapse Moreneuro-trans-mitter Lessneuro-trans-mitter –50 –50 Axon –50 Receptor potential –70 –70 –70 Membranepotential (mV) Membranepotential (mV) Membranepotential (mV) Action potentials 0 0 0 –70 –70 –70 5 6 7 1 2 3 4 0 0 0 1 2 3 4 5 6 7 1 2 3 4 5 6 7 Time (sec) Time (sec) Time (sec) (b) Vertebrate hair cells have specialized cilia or microvilli (“hairs”) that bend when sur-rounding fluid moves. Each hair cell releases an excitatory neurotransmitter at a synapse with a sensory neuron, which conducts action potentials to the CNS. Bending in one direction depolarizes the hair cell, causing it to release more neurotransmitter and increasing frequency Figure 49.2b

12. Electromagnetic Receptors • Electromagnetic receptors detect various forms of electromagnetic energy • Such as visible light, electricity, and magnetism

14. Diverse photoreceptors • Eye cups • Compound Eyes • Single-lens eyes (used by vertebrates)

16. Human retina • Rods are sensitive to light but do not distinguish colors • Cones distinguish colors but are not sensitive

19. Rod Cells are light sensitive but do not distinguish colors • Cone Cells are not as light sensitive as rods but provide color vision and are mostly concentrated on the fovea

20. The absorption of light by rhodopsin initiates a signal-transduction pathway

21. Cones • Have 3 subclasses and more complex than the rhodopsin mechanism. • Each has its own type of photopsin.

22. Retina, cerebral cortex and processing visual info • Visual processing begins with rods and cones synapsing with bipolar cells which then synapse with ganglion cells. • Visual processing in the retina also involves horizontal cells and amacrine cells.

24. Lateral pathways involving horizontal or amacrine cells can inhibit adjacent pathways: • Photoreceptors  horizontal cells other photoreceptors • Photoreceptorsbipolar cells amacrine cells ganglion cells • The resulting lateral inhibition (More distance photoreceptors and bipolar cells are inhibited  sharpens edges and enhances contrast in the image)

25. The optic nerves of the two eyes meet at the optic chiasm. • Ganglion cell axons make up the optic tract. • Most synapse in the lateral geniculate nuclei of the thalamus • The neurons then convey information to the primary visual cortex of the optic lobe

27. Cochlea Stapes Axons ofsensoryneurons Oval window Vestibularcanal Perilymph Apex Base Roundwindow Tympaniccanal Basilar membrane • Vibrations create pressure waves in the fluid in the cochlea that travel through the vestibular canal and strike the round window

28. Cochlea(uncoiled) Apex(wide and flexible) Basilarmembrane 500 Hz(low pitch) 1 kHz 2 kHz 4 kHz 8 kHz Frequency producing maximum vibration 16 kHz(high pitch) Base(narrow and stiff)

29. Ear and Balance • Behind the oval window is a vestibule that contains the utricle and saccule • The utricle opens into three semicircular canals

30. Utricle and saccule respond to changes in head position relative to gravity and movement in one direction. • Hair cells are projected into a gelatinous material containing otoliths. • Semicircular canals respond to rotational movements of the head

32. Most fish and amphibians have a lateral line system along both sides of their body • Provides a fish with information concerning its movement through water or the direction and velocity of water flowing over its body.

33. Invertebrates with gravity sensors and sound that are sound-sensitive • Statocysts are mechanoreceptors that function in an invertebrates sense of equilibrium

34. Insects • Sound sensitivity in insects depends on body hairs that vibrate in response to sound waves. • Many insects have a tympanic membrane stretched over a hollow chamber

37. Taste pore Sugar molecule Taste bud Sensoryreceptorcells Sensoryneuron Tongue Sugar Adenylyl cyclase G protein Sugarreceptor ATP cAMP Proteinkinase A SENSORYRECEPTORCELL K+ Synapticvesicle —Ca2+ Neurotransmitter Sensory neuron Transduction in taste receptors

38. Mammals’ olfactory receptors line the upper portion of the nasal cavity • The binding of odor molecules to olfactory receptors initiate signal transduction pathways involving a G-protein-signaling pathway and, often, adenylyl cyclase and cyclic AMP.

39. The key to flight is the aerodynamic structure of wings

40. Peristaltic motion pushes against hydrostatic skeleton

49. When myosin binds to actin it forms an ATPase which releases the energy from ATP for contraction