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Leaves. Leaf Structure. Epidermal tissue system chloroplasts only in guard cells Ground tissue system mesophyll Vascular tissue system Leaf adaptations sun and shade leaves xerophytes. CO 2. sunlight, water. Dark reacti ons (Calvin cycle). Light reactions. Oxygen (O 2 ).
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Leaf Structure • Epidermal tissue system • chloroplasts only in guard cells • Ground tissue system • mesophyll • Vascular tissue system • Leaf adaptations • sun and shade leaves • xerophytes
CO2 sunlight, water Dark reacti ons (Calvin cycle) Light reactions Oxygen (O2) Glucose (C6H12O6) (carbohydrates) Photosynthesis
Start with the Big Picture • It’s all about energy • potential vs kinetic energy • Nearly all energy used on earth comes from the sun • Nearly all energy used on earth ends up as heat • Metabolism • the way living things manage energy
Laws of Thermodynamics • Energy can neither be created nor destroyed but it can be converted from one form to another • In any transfer of energy there is always loss of useful energy, usually as heat
What makes chemical reactions happen? • Chemical bonds have potential energy • When the bonds are broken, the energy is released • Cells transfer energy from one molecule to another, often as a proton (H+) or electron (e-) • Enzymes are proteins that speed up chemical reactions • ATP has high-energy phosphate bonds • ATP bonds come from glucose • Glucose comes from...
Solar energy captured in chemical bonds Chemical bond energy is released to do work Metabolism summary • Photosynthesis • light reactions • dark reactions • Respiration • glycolysis • Krebs cycle • electron transport • fermentation
CO2 sunlight, water Dark reacti ons (Calvin cycle) Thylakoid membrane ATP, NADPH Light reactions Chlo roplast stroma oxygen Glucose (C6H12O6) (carbohydrates) Photosynthesis
Light Reactions of Photosynthesis • Chlorophyll is a pigment in the thylakoid membrane • It can absorb light energy • The absorbed energy knocks an electron out of the chlorophyll molecule • This electron is transported from one molecule to another • The energy is used to make ATP • The original electron is replaced with one from a molecule of water, releasing O2 Movie 1 Movie 2
Dark Reactions (Calvin cycle) • Occurs in the stroma of the chloroplast • CO2 diffuses into spongy mesophyll cells • RUBISCO is the enzyme that fixes the CO2 into a carbon compound • it’s the most abundant protein on earth • a very complicated enzyme!
Calvin cycle Movie
The atmosphere contains oxygen because of photosynthesis
Oxygen causes a problem for RUBISCO • RUBISCO binds O2 as well as CO2 • the substrates compete for the active site • O2 results in no new sugar • Someplants have evolved a way to avoid this wasteful reaction
ATP <
Water molecules < Waterbox
Companioncells Sieve cells <
Dicot leaf surface Monocot leaf surface
Palisade parenchyma Upper epidermis Xylem Stoma Phloem Spongy parenchyma Lower epidermis
Sun leaf and shade leaf sections Genetic vs environmental differences in leaves
Leaf cross section Which tissue holds the water?
Parallel venation (monocots) Net or pinnate venation (dicots)
Typical horizontal leaf Typical vertical leaf