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Chapter 43 The Biosphere (Sections 43.5 - 43.9). 43.5 Deserts. Deserts receive less than 10 centimeters (4 inches) of rain per year Most are located at about 30° north and south latitude, or in rain shadows desert
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43.5 Deserts • Desertsreceive less than 10 centimeters (4 inches) of rain per year • Most are located at about 30° north and south latitude, or in rain shadows • desert • Biome with little rain and low humidity; plants that have water-storing and conserving adaptations predominate
Desert Conditions • Deserts tend to have low humidity, large daily temperature shifts, and little topsoil
Desert Conditions O horizon: Pebbles, little organic matter A horizon: Shallow, poor soil B horizon: Evaporation causes salt buildup; leaching removes nutrients C horizon: Rock fragments from uplands Fig. 43.10, p. 730
Adaptations to Desert Conditions • Many desert plants have adaptations, such as spines, that reduce water loss and deter herbivory • Some desert plants, such as cacti, store water during the wet season, for use in drier times • Woody desert shrubs such as mesquite and creosote have extensive, efficient root systems
Adaptations to Desert (cont.) • Diversity is highest in regions where moisture is available in more than one season • CAM plants (cactuses, agaves, euphorbs) conserve water by opening stomata only at night when temperature declines • Annuals have a life cycle that allows them to sprout and reproduce in the short time that the soil is moist
Animal Adaptations • Some animals, such as the desert kangaroo rat, minimize water loss with highly efficient kidneys • Most desert animals, including bats, are not active at the height of daytime heat • The Sonoran desert tortoise hibernates during the cold winter, when little food is available
The Crust Community • In many deserts, the soil is covered by a desert crust (including cyanobacteria, lichens, mosses, and fungi) that holds soil in place • Cyanobacteria also fixes nitrogen and makes ammonia available for plants
43.6 Grasslands • Grasslandsform in the interior of continents between deserts and temperate forests • Perennial grasses adapted to fire, grazing , strong winds and infrequent rain, are the main plants in grasslands • grassland • Biome in the interior of continents where grasses and nonwoody plants adapted to grazing and fire predominate
Temperate Grasslands • Temperate grasslands are warm in summer, cold in winter, with rains throughout the year • Grass roots help hold thick topsoil in place, preventing erosion by constant winds
Temperate Grasslands A horizon: Alkaline, deep, rich in humus B horizon: Percolating water enriches layer with calcium carbonates A Prairie soil profile. Fig. 43.15a, p. 732
North American Grasslands • North America’s temperate grasslands are shortgrass and tallgrass prairies • Much of the American Great Plains shortgrass prairie was plowed in the 1930s, turning the region into a “Dust Bowl” • Nearly all tallgrass prairie has been converted to cropland
Savannas • Savannas are broad belts of grasslands with scattered shrubs and trees, that lie between tropical forests and deserts • Temperatures are warm year-round, but rainfall is seasonal • African savanna supports herds of hoofed grazers (such as wildebeest) and predators that feed on them (such as lions)
43.7 Dry Shrublands and Woodlands • Rains occur seasonally in dry shrublands, and lightning-sparked fires sometimes sweep through shrublands during the dry season • Dry shrublands in California are known as chaparral • chaparral • Biome of dry shrubland in regions with hot, dry summers and cool, rainy winters
Adaptations to Chaparral • Foliage of many chaparral shrubs has oils that deter herbivores and also make the plant highly flammable • Chaparral plants are adapted to occasional fires: • Some grow back from root crowns after a fire • Some have seeds that germinate only after they are exposed to heat or smoke
California Chaparral • In California, where homes are often built near chaparral, fires frequently cause property damage
Dry Woodlands • Dry woodlands prevail where the seasonal rainfall is slightly higher than in dry shrublands • Examples: Eucalyptus forests of Australia, and California’s oak forests
43.8 Broadleaf Forests • Broadleaf (angiosperm) trees are the main plants in semi-evergreen forests, and in temperate and tropical deciduous forests • Semi-evergreen forests occur in the tropics of Southeast Asia and India, and include broadleaf (angiosperm) trees that retain leaves year-round, and deciduous broadleaf trees • In semi-evergreen forests, deciduous trees shed their leaves at the start of the dry season
Temperate Deciduous Forests • A deciduous plant sheds leaves annually, prior to a season when cold or dry conditions would not favor growth • Temperate deciduous forests are found in parts of eastern North America, western/central Europe, and parts of Asia • temperate deciduous forest • Northern Hemisphere biome in which the main plants are broadleaf trees that lose their leaves in fall and become dormant during cold winters
Temperate Deciduous Forests (cont.) • Temperate deciduous forests form where 50 to 150 centimeters (20–60 inches) of precipitation falls throughout the year • Winters are cool and summers are warm • Leaves often turn color before dropping in autumn • Trees remain dormant in winter, flower and put out new leaves in spring
Temperate Deciduous Forests (cont.) • Rich soil and a somewhat open canopy lets sunlight through and allows understory plants to flourish
Temperate Deciduous Forests (cont.) O horizon: Scattered litter A horizon: Rich in organic matter above humus layer unmixed with minerals B horizon: Accumulated minerals leached from above C horizon: Poorly weathered rocks Fig. 43.19b, p. 734
Temperate Deciduous Forests (cont.) • Different tree species (such as oak, or beech and maple) characterize different regions of these forests • Animals in North American deciduous forests include grazing deer, seed-eating squirrels and chipmunks, and omnivores (raccoons, opossums, black bears)
Tropical Rain Forests • Tropical rain forestsform between latitudes 10° north and south in Africa, East Indies, Asia, South and Central America • 130 to 200 centimeters (50 to 80 inches) of rain falls throughout the year • tropical rain forest • Highly productive and species-rich biome in which year-round rains and warmth support continuous growth of evergreen broadleaf trees
Tropical Rain Forests (cont.) • Tropical rain forest is the most productive, structurally complex, and species-rich biome • Photosynthesis occurs year-round; but trees often form a closed canopy that blocks light from the forest floor • Deforestation is an ongoing threat to tropical rain forests in developing countries with fast-growing human populations
Tropical Rain Forests (cont.) • Decomposition and mineral cycling happen fast in the warm, moist environment • Soils are highly weathered, heavily leached, and are very poor nutrient reservoirs
Tropical Rain Forests (cont.) O horizon: Sparse litter A–E horizons: Continually leached; iron, aluminum left behind impart red color to acidic soil B horizon: Clays with silicates, other residues of weathering Fig. 43.20b, p. 735
43.9 Coniferous Forests • Conifers withstand harsher conditions than broadleaf trees, so they grow farther north and at higher altitudes • The coniferous forest that extends across northern Asia, Europe, and North America – known as boreal forest or taiga– is the most extensive land biome • boreal forest • Extensive high-latitude forest of the Northern Hemisphere; conifers are the predominant vegetation
Coniferous Forests (cont.) • Conifers (evergreen trees with seed-bearing cones) such as pine, fir, and spruce are the main plants in coniferous forests • Conifers are adapted to conserve water during drought or times when the ground is frozen – winters are long, cold, and dry; and most rain falls in the cool summer • Moose are the dominant grazers
Other Coniferous Forests • Montane coniferous forests of spruce, fir, and pine extend southward through great mountain ranges • Spruce and fir dominate highest elevations • Firs and pines occupy lower elevations • Coniferous forests in temperate lowlands along the Pacific coast from Alaska into northern California hold the world’s tallest trees: • Sitka spruce to the north • Coast redwoods to the south
Other Coniferous Forests (cont.) • New Jersey pine barrens are a mixed forest of pitch pines and scrub oaks that grow in sandy, acidic soil • Southeastern pine forests hold fast-growing loblolly pines that survive periodic fires that kill most hardwood species – these pines are a major source of lumber and wood pulp