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Succession in Reefs. 4 stages : Pioneering or Stabilization a. Accumulation of skeletal debris b. Creation of a surface with some relief. Succession in Reefs. 2. Colonization a. Branching & encrusting organisms b. Low diversity. Succession in Reefs. Diversification
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Succession in Reefs • 4 stages : • Pioneering or Stabilization a. Accumulation of skeletal debris b. Creation of a surface with some relief
Succession in Reefs 2. Colonization a. Branching & encrusting organisms b. Low diversity
Succession in Reefs • Diversification a. Increased diversity of organisms b. Takes the form of a massive structure
Succession in Reefs • Domination a. “lamellar framestones” b. Evidence of high water energy (rudstones) c. Species diversity is reduced
Reef Evolution www.netrocks.net inyo.topcities.com Stromatolites 12 bybp
Reef Evolution Back to “pond slime” Loss of “ecospace” What happens with a sea-level fall? 12 bybp
The Ordovician www.ammonite.free-online.co.uk
Mammelon Astrorhizae Stromatoporoids
Ordovician Reefs (505-438 my) Bryozoan Stromatoporoid Peter Sholle Pratt & James (1982)
Early Silurian Extinction
REEF DIVERSIFICATION Sea-level is back up
The Silurian World 20° OCEAN Equator 20° Chris Scotese
Silurian Time430 mybp Equator Reefs Scotese (1997) Dott & Prothro (1994)
New Life in the Silurian Corals Through Time 0 Wider = more abundant 200 400 600
Crinoids Karla Parsons-Hubbard (Oberlin College) Beloit College Collection
40 million years later.. Opening up; Sea-level rise Shallow, clear water
Devonian 12 bybp
Late Devonian Why??
Reefs made of sponges….. The Capitan Reef Complex Sea Level Back Up Sea-Level Drops Just when things were looking good….. Devonian
Photo by Peter Sholle Photo by Peter Sholle Photo by Mark Eberle
Extinction Why? • Sea level - no • Climate change - maybe • Ocean chemistry? Reefs disappear!!!!!!!!
A watershed event in reef history?? Zooxanthellae 20 million yr gap
Watch here… Chris Scotese
The cup shaped form is back….. • but it’s a mollusk this time??? A broadening tropics An open pan-equatorial seaway
Open tropical seaway No glaciers !
High temperatures “Greenhouse” climate much warmer than today Too warm for corals Why mollusks and not corals???
Cretaceous Reef Line Reefs Today
Rudist Reefs They photosynthesized?? the missing article…… Photo by D. Schumann (from Woods, 1999) Drawing by John Sibbick (in Woods, 1999)
Dewey McLean Extinction The End of the Cretaceous Images by Mary Parrish, NMNH Irridium Cretaceous Time
Major extinction 95% of species disappear
Reefs have stayed pretty much the same ever since Rest of the Cenozoic
Cambrian “Explosion” Mostly non-reef forms
Increased Calcification Jaws -->BioE Calcification
“Party Crashers” Faster Growth Environment Sea Level More Complex Increased grazing No O2 Wilson Ramirez