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Populations. Populations. Population - a group of organisms of the same species which have the potential to interbreed – or a population is a group of organisms of the same species occupying a particular place at a particular time
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Populations • Population - a group of organisms of the same species which have the potential to interbreed – or a population is a group of organisms of the same species occupying a particular place at a particular time • Populations have a number of properties which are not possessed by individual organisms - this is because a population is the sum of many organisms interacting
Starling Murmuration http://vimeo.com/31158841
Properties dealing with changes in population size • Natality - may think of this as births, but includes more than just birth - hatching, germination, fission • Natality includes idea of fecundity - number of offspring produced per unit time - we are most concerned with realized fecundity - actual number of survivors • Mortality - death rate - its converse is survivorship - mortality looks at how many die per unit time, survivorship at how many don't die per unit time • Longevity examines life-span of individuals - again we are most interested in realized longevity, not potential longevity • Immigration - individuals moving into a population • Emigration - individuals leaving a population
What is an individual? • unitary organism - individuals are highly determinate in form and while growing pass through predictable (innately determined) sequences of life history stages • modular organisms - zygote develops into unit, or module, which produces more modules thus producing an organism with a variable number of modules, whose development is unpredictable and strongly influenced by environmental factors
A classic modular organism Bryozoan colony
Genets and Ramets • ramet - a module with the potential for a separate existence • genet - the "genetic individual"; the collection of all modules derived from a single zygote
Sampling to collect population data Census - most basic sampling - count and determine age of all individuals in population, count again later Several ways to subsample and estimate population size: 1. Determine total area in which population occurs, count all individuals in small plots, multiply average number in plots to get total, repeat at later dates - works best for sessile organisms 2. Mark-recapture methods 3. Catch per unit effort 4. Miscellaneous methods – traps, counts of fecal pellets, counts of vocalizations, feeding damage on plants, radar counts, roadside sightings, fur or pelt records, roadkill
Metapopulations • A metapopulation is a series of small, separate populations united together by dispersal
Bay Checkerspot Jasper Ridge
Demography • Demography is the study of processes that influence population size - it is the way we study changes brought about by births, deaths and dispersal For Northwest Montana, Greater Yellowstone, Central Idaho
The Fundamental Equation of Ecology – Harper 1977 Δ N = B – D + I – E Change in Number = Births – Deaths + Immigration - Emigration John L. Harper – 1925-2009
Rearrange Fundamental Equation Δ N = B – D + I – E Nfuture = Nnow + B – D + I – E