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GROWTH RESPONSES AND BIOLOGICAL RHYTHMS IN PLANTS CH 33. Tropisms orient plant growth toward or away from environmental stimuli Plants sense and respond to environmental changes In a variety of ways. Light. Tip covered by opaque cap. Base covered by opaque shield. Tip separated
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GROWTH RESPONSES AND BIOLOGICAL RHYTHMS IN PLANTSCH 33 • Tropisms orient plant growth toward or away from environmental stimuli • Plants sense and respond to environmental changes • In a variety of ways
Light Tip covered by opaque cap Base covered by opaque shield Tip separated by mica Tip separated by gelatin block Tip covered by trans- parent cap Tip removed Control Darwin and Darwin (1880) Boysen-Jensen (1913) • Response to Light • Phototropism- bending in response to light • May result from auxin moving from the light side to the dark side of a stem
Response to Gravity • A response to gravity, orGravitropism • May be caused by the settling of special organelles on the low sides of shoots and roots • This may trigger a change in the distribution of hormones Figure 33.9A
Response to Touch • Thigmotropism, a response to touch • Is responsible for the coiling of tendrils and vines around objects Figure 33.9B
Plants mark the seasons by measuring photoperiod • Photoperiod • Is the relative lengths of night and day • The timing of flowering • Is one of the seasonal responses to photoperiod
24 Darkness Critical night length Flash of light Time (hr) Light 0 Long-day (short-night) plants Short-day (long-night) plants • Plants whose flowering is triggered by photoperiod fall into two groups • Short-day (long-night) plants • Long-day (short-night) plants Figure 33.11
Phytochromeis a light detector that may help set the biological clock • Phytochromes are proteins with a light-absorbing component • That may help plants set their biological clock and monitor photoperiod
24 Darkness Critical night length Flash of light Time (hr) Light 0 Long-day (short-night) plants Short-day (long-night) plants Results
4 5 Recruitment of wasp Wasp lays eggs 3 Synthesis and release of chemical attractants 1 2 Damage to plant and chemical in caterpillar saliva Signal transduction pathway • Defenses Against Herbivores • Some plants recruit predatory animals • To help defend them against certain herbivores Adapted from Edward Farmer, “Plant Biology: New Fatty Acid–Based Signals: A Lesson from the Plant World” Science 276 (1997), p. 912. 1997 American Association for the Advancement of Science. Plant cell Figure 33.14A
PLANT DEFENSES • Defenses against herbivores and infectious microbes have evolved in plants • Plants use chemicals • To defend themselves from both herbivores and pathogens
Local defenses include • Microbe-killing chemicals and sealing off the infected area • Hormones • Trigger generalized defense responses in other organs (systemic acquired resistance)