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Explore the classification and relationships of organisms through taxonomy and evolutionary systematics. Learn about binomial nomenclature and molecular taxonomy to understand biodiversity and phylogeny. Discover the significance of monophyletic and paraphyletic taxa.
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Chapter 22 Understanding Diversity:Systematics
Systematics • Diversity of organisms • Evolutionary relationships • Biodiversity • The variety of organisms • The ecosystems they form
Taxonomy • Naming • Describing • Classifying • Classification • Grouping organisms by their similarities or relationships
Binomial nomenclature • Genus + specific epithet • Homo sapiens • Quercus alba • Linnaeus, mid-18th century • Facilitates international science
Taxonomic categories • Kingdom • Phylum • Class • Order • Family • Genus • Species
Six Kingdoms • Based on cell structure & nutrition • Archaebacteria • Eubacteria • Protista • Fungi • Plantae • Animalia
Three domains • Based on molecular differences • Archae • Lack simple RNA polymerase • Eubacteria • Peptidoglycan in cell walls • Eukarya • Eukaryotes
Phylogeny • Evolutionary history of a group of organisms from a common ancestor • Classifications based on common ancestors • Traditionally based on structural similarities
Homology • Structure derived from a recent common ancestor • Wing bones in birds and bats • Homoplasy • Superficially homologous but independently acquired • Body form in sharks and dolphins
Plesiomorphic characters • Shared ancestral characters • Distant common ancestor • Synapomorphic characters • Shared derived characters • More recent common ancestor
Molecular systematics • Determines evolutionary relationships by comparing macromolecules • Ribosomal RNAs 5S, 16S, 23S • Transcribed from highly conserved DNA regions • Mitochondrial DNA
Molecular clocks • Measure time since divergence from a common ancestor by the number of differences in nucleotide or protein sequences • Rate of change must be constant
Monophyletic taxon • All descendants of most recent common ancestor • Mammals • True evolutionary relationships • Clade
Paraphyletic taxon • Common ancestor but not all descendants • Class Reptilia does not include birds even though they share a recent common ancestor • Avoided by cladistic systematists
Polyphyletic group • Does not share recent common ancestors • Protista • Shares homoplastic features • Misrepresents evolutionary relationships
Evolutionary systematics • Evolutionary branching • Extent of divergence • Structural and other changes • Combination of shared ancestral characters and shared derived characters
Cladistics • Emphasizes common ancestry over phenotypic similarity • Only shared derived characters • Cladograms • Parsimony guides choices
Outgroup analysis • Separate ancestral and derived characters • Outgroup taxon diverged earlier than others • Outgroup represents the ancestral condition