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Evolution of Populations. 2010. The Modern Synthesis. Population genetics integrates Darwinian evolution and Mendelian Genetics Important terms in population genetics: Population: group of individuals of same species
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The Modern Synthesis • Population genetics integrates Darwinian evolution and Mendelian Genetics • Important terms in population genetics: Population: group of individuals of same species Species: groups of populations that are capable of interbreeding and producing fertile offspring in nature Gene pool: total aggregate of genes in a population
Hardy – Weinberg Theorem • Gene pool of non-evolving population • States that the frequency of alleles and genotypes in a population’s gene pool remain constant unless acted upon by agents other than Mendelian segregation and recombination of alleles. p+q = 1 Hardy Weinberg Equilibrium: p2 + 2pq + q2 = 1 P2 = Frequency of RR phenotype 2pq -= Frequency of Rr phenotype q2= Frequency of rr phenotype
Conditions that must be met: Very large population size. No migration No net mutations Random mating No natural selection Do you that any natural populations are in Hardy-Weinberg equilibrium?
Microevolution • Generation to generation change in the population’s frequency of alleles • 2 main causes: genetic drift – change due to chance, small populations & natural selection • Other causes -
Bottleneck Effect • Drastically reduced population sizes • Small population size may not be representative of original gene pool • Reduction in genetic variablility
The Founder Effect Occurs when a few individuals colonize an isolated habitat from a larger population
Gene Flow • Genetic exchange through migration – gain or loss of alleles
Mutation Substitutes one allele for another
Speciation – Biological Species Concept • Prezygotic Barriers: habitat isolation, behavioral isolation, temporal isolation, mechanical isolation, gametic isolation • Postzygotic Barriers: reduced hybrid viability, reduced hybrid fertility, hybrid breakdown
Types of Speciation • Allopatric: geographic barriers, disruption of gene flow • Sympatric: genetic causes – polyploid (plants), other genetic factors
Tempo of Evolution • Darwin – gradual continual rate • Punctuated equilibrium model: spurts of rapid change followed by periods of relatively little change (Ex. Species exists for 5 million years but most morphological changes in 50,000 yrs – just 1%).
Phylogenies • Many lines of evidence compiled • Biogeographical, fossil, morphological similarities (homologous not analogous structures), genetic evidence
Categories of Evolution • Convergent evolution - acquisition of the same biological trait in unrelated lineages. • Leads to analogous structures
Parallel evolution is the development of a similar trait in related, but distinct, species descending from the same ancestor, but from different clades
Divergent evolution - accumulation of differences between groups which can lead to the formation of new species, usually a result of diffusion of the same species adapting to different environments
Major Lineages of Life 3 domains: Bacteria, Eukaryota, Arachae