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EAR. EAR. External (outer) ear - channels sound waves to middle ear Middle ear - directs sound waves to oval window Internal (inner) ear - contains mechanoreceptors (hair cells) for hearing & equilibrium. Outer Ear.
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EAR • External (outer) ear - channels sound waves to middle ear • Middle ear - directs sound waves to oval window • Internal (inner) ear - contains mechanoreceptors (hair cells) for hearing & equilibrium
Outer Ear • Fleshy auricle (pinna) directing air vibrations down auditory canal (external auditory meatus)
MIDDLE EAR • Air-filled cavity in temporal bone • Separated from the exterior by the tympanic membrane (ear drum) • Filled with air by auditory tube (Eustachian tube) connected to nasopharynx • The auditory tube opens during swallowing or yawning to equalize air pressure on both sides of eardrum. • Ear ossicles span tympanic cavity - held in place by tensor tympani and stapedius muscles
INNER EAR • Consists of: 1. Bony labyrinth - passageways in the temporal bone that contain perilymph 2. Membranous labyrinth - fleshy tubes lining bony tunnels filled with endolymph and floating in perilymph • Houses cochlea and vestibular apparatus
Anatomy of the Cochlea 2.5 coils 3 fluid-filled chambers Organ of Corti
HOW THE EAR WORKS • Eardrum vibrates & creates enough force/unit area at oval window to vibrate the endolymph in the scala vestibuli. Vibrations excite hair cells. • Contraction of tensor tympani & stapedius protect the cochlea in response to loud noises (tympanic reflex) • Contraction of tensor tympani pulls eardrum inward and tightens it. • Contraction of the stapedius reduces mobility of stapes. • The ear muscles also contract during speaking so we can hear others.
SENSORY CODING • Loudness produces vigorous vibrations that excite many hair cells. This triggers higher frequency of action potentials that the brain interprets as loudness. • Determination of pitch depends on which part of the basilar membrane vibrates in the organ of Corti.
Frequency Response of Basilar Membrane Peak amplitude of wave varies with frequency
EQUILIBRIUM • The vestibular apparatus is concerned with equilibrium. • Maculae in saccule & utricle perceive head orientation & maintain static equilibrium • Maculae & cristae maintain dynamic equilibrium. • Maculaeperceivelinear acceleration • Cristae of semicircular ducts perceive angular acceleration
SACCULE & UTRICLE • Contain maculae • A macula is a patch of hair cells with stereocilia & one kinocilium buried in a gelatinous otolithic membrane weighted with granules called otoliths.
Crista Ampullaris of Semicircular Ducts • A crista ampullaris consists of hair cells buried in a mound of gelatinous membrane. • The orientation of one duct causes other ducts to be stimulated.
Crista Ampullaris & Head Rotation • As the head turns, endolymph lags behind pushing cupulae and stimulating hair cells
THE VESTIBULAR NERVE • Hair cells of macula sacculi, macula utriculi & semicircular ducts synapse on the vestibular nerve. • Fibers end in the vestibular nucleus of pons, cerebellum, nuclei of cranial nerves controlling eye, head and neck movements.