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Intro to Lab #9 / Chpt. 36 Plant Structure and Transport pg. 744 - 753. Transport in Plants. Did you know, an average size maple tree looses 200 L of water per hour during the summer via. transpiration. What would happen if this water were not replaced by the roots???. Question ?.
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Intro to Lab #9 / Chpt. 36 Plant Structure and Transport pg. 744 - 753
Transport in Plants Did you know, an average size maple tree looses 200 L of water perhour during the summer via. transpiration. What would happen if this water were not replaced by the roots???
Question ? • How do plants move materials from one organ to the other ?
TRANSPIRATION TRANSPIRATION - when a plant looses water vapor from the inside of the leaf, to the outside environment -via. stomata
COHESION - TENSIONTHEORY - water is pulled up from roots to leaves via. higher in the leaf to a lower in the air
Water & mineral absorption to Xylem: • Water absorption from soil - osmosis - aquaporins
ROOT HAIRS = increase surface area for the absorption of water & minerals
Water & mineral absorption to Xylem: Mineral absorption • active transport • proton pumps • active transport of H+ proton pumps
Water & mineral absorption to Xylem: • Proton pumps • active transport of H+ ions out of cell • chemiosmosis • H+ gradient • creates membranepotential • difference in charge • drives cation uptake • creates gradient • cotransport of othersolutes against theirgradient proton pumps
Transport in Plants uptake and loss by individual cells: The survival of plant cells depends on the ability to balance the uptake & loss of H2O
Remember this? *Presence of the cell wall adds physical pressure, this affects movement into the cell negatively!
Remember this? *Solute concentration also affects movement:
Transport in Plants How can we predict the direction of osmosis when a plant cell is surrounded by a solution???
Transport in Plants WATER POTENTIAL
Remember this? Water moves from the solution of higherwaterpotential, to a solution of lowerwaterpotential. For plants, it is not just enough to know if the extracellular solution is hypotonic or hypertonic.
Remember this? *Measured in MEGAPASCALS • Has two components: • Pressure potential: yr • Solute potential: yp • y = yr + yp
Remember this? You have to know these two numbers first, when determining water potential. You must know water potential in order to know which direction water will move! Pressure potential: yrSolute potential: yp y = yr + yp
Remember this? any solution will have a negative w.p. adding solutes, lowers w.p.
Transport in Plants *Solute concentration affects movement
Transport in Plants *External pressure on a solution counters its tendency to take up water due to the presence of solutes
Transport in Plants = + P S
Transport in Plants the force that moves water across the membranes of plant cells =
COHESION - TENSIONTHEORY - water is “pulled” b/c of it’s cohesiveproperty
Evolutionary advantage of ROOT HAIRS: add surface area - thus increasing amount of H20/mineral solution coming into the plant!!
Mycorrhizae increase absorption • Symbiotic relationship between fungi & plant • symbiotic fungi greatly increases surface area for absorption of water & minerals • increases volume of soil reached to plant • increases transport to host plant
COHESION - TENSIONTHEORY - Because higher water potential outside
COHESION - TENSIONTHEORY - minerals absorbed from soil into root~ creates “rootpressure” that pushes H2O/soil solution into xylem
COHESION - TENSIONTHEORY - • uptake of soil solution by root hairs to apoplasticroute-through the cell walls
COHESION - TENSIONTHEORY - • uptake of minerals & water by root hairs to symplasticroute - through plasmodesmata
Controlling the route of water in root • Endodermis • cell layer surrounding vascular cylinder of root • lined with impermeable Casparian strip • forces fluid through selective cell membrane • filtered & forced into xylem cells
Controlling the route of water in root • some soil solution from apoplast route, diffuses into the symplastic route -through the plasma membrane
COHESION - TENSIONTHEORY - Water then travels thru the root where the Casparian strip forces H2O into the xylem REMEMBER: the casparian strip blocks water from entering via. The apoplastic route
COHESION - TENSIONTHEORY - • only minerals in the symplastic route can detour around the Casparian Strip AND PASS INTO THE VASCULAR XYLEM!!
Root anatomy eudicot monocot
COHESION - TENSIONTHEORY - • XYLEM vessels transport water and minerals upward into the shoot system (stem and veins)
COHESION - TENSIONTHEORY - problem… this can only take the water “so far”
COHESION - TENSIONTHEORY - At the leaf of the plant, water loss due to evaporation is taking place.