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This article discusses the historical non-spatial approach to ecology and highlights the need to consider spatial factors. It explores the Vancouver Island marmot as a case study, examining natural tree succession, forestry disturbance, climate, and prey-predator dynamics. The article also delves into the history and development of landscape ecology as a science, emphasizing the importance of studying large-scale landscapes and considering human impacts. It further discusses the emergence of landscape ecology as a dynamic field, exploring concepts of scale, hierarchy, and the role of disturbance in ecosystem response. Advances in technology and the four key objectives of landscape ecology are also explored, along with the importance of this field in conservation and management efforts.
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Space Is the Final Frontier in Ecology • History of ecology = largely nonspatial e.g., *competitors mixed perfectly with prey *homogeneous ecosystems with uniform distributions of resources • Simple population models
Space Is the Final Frontier in Ecology • History of ecology = largely nonspatial e.g., *competitors mixed perfectly with prey *homogeneous ecosystems with uniform distributions of resources • But ecology = fundamentally spatial • ecology = interaction of organisms with their [spatial] environment
Vancouver Island marmot(Marmota vancouverensis) ~100 left Isolated from hoary and Olympic marmots
Vancouver Island marmot(Marmota vancouverensis) Natural tree succession
Vancouver Island marmot(Marmota vancouverensis) • Logging – disjunct patches • max. dispersal = 7 km • - Forestry disturbance • Climate • Prey-Predator Dynamics
History of Landscape Ecology • “European School” • roots back to those of ecology • typing, classifying, naming • landscape architecture, planning, designing • “American School” • young = early 1980’s • focus on natural systems • theory & models, some field experiments
Landscape Ecology as a Science? • Meta-analysis of papers in Landscape Ecology (Wiens 1992) – 1st 5 yrs of journal • Most studies are large scale (landscapes are big) • Most studies are descriptive or conceptual • Experiments difficult to conduct, thus modeling • Emphasis on vegetation pattern / land use pattern • Humans = impt. part of systems
Landscape Ecology as a Science? • Hobbs (1997) – 2nd 5 years of Landscape Ecology • Less descriptive studies • More “methods” and modeling studies, no expts. • More quantitative / statistical analysis (spatial stats)
Structure Function Emergence of Landscape Ecology ? Equilibrium View • Constant species composition • Disturbance & succession = subordinate factors • Ecosystems self-contained • Internal dynamics shape trajectory • No need to look outside boundaries to understand ecosystem dynamics ? ? ?
Structure Function Emergence of Landscape Ecology Dynamic View • Disturbance & ecosystem response = key factors • Disturbance counter equilibrium • Ecosystems NOT self-contained • Multiple scales of processes, outside & inside • Essential to examine spatial & temporal context
Emergence of Landscape Ecology Broad-scale Environmental Issues Landscape Ecology Concepts of Scale/Hiearchy Dynamic View Advances in Technology
Landscape ecology focuses on spatial heterogeneity and pattern. 1) detecting pattern 2) define origin of pattern 3) predicting change 4) relating pattern to processes & mgt
1 m2 • Often, spatial extents much larger than in traditional ecology… • But, emphasize spatial pattern at relevant scale 1 km2 10’s km2
Often, focus on role of humans in creating and influencing patterns… • But, humans only one (albeit dominant) agent
Importance of Landscape Ecology to Conservation & Management Ecosystem context matters Ecosystem function depends on interplay of pattern & process Policy & legal mandate to adopt landscape perspective
Importance of Landscape Ecology to Conservation & Management • Ecosystem context matters Metapopulations
Importance of Landscape Ecology to Conservation & Management • Ecosystem function depends on interplay of pattern & process Habitat fragmentation & Modified Disturbance Regimes
Importance of Landscape Ecology to Conservation & Management • Policy & legal mandate to adopt landscape perspective