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Phloem - II. Primary Phloem. Protophloem Metaphloem. Metaphloem Sunflower and Corn. Secondary Phloem in Dicots. Sieve tube members Companion cells Fibers Sclereids Regular parenchyma Axial parenchyma Ray parenchyma Dilatation tissue – axial or ray. Secondary phloem in pumpkin stem.
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Primary Phloem • Protophloem • Metaphloem
Secondary Phloem in Dicots • Sieve tube members • Companion cells • Fibers • Sclereids • Regular parenchyma • Axial parenchyma • Ray parenchyma • Dilatation tissue – axial or ray
Secondary phloem in pumpkin stem 1 = STM; 2 = Sieve plate; 3 = Companion cell; 4 = Vascular cambium 5 = Vessel elements
Secondary phloem is short lived • As new secondary phloem is produced, older STM and CC are crushed and may be obliterated • This is caused by the expansion of the stem diameter during secondary growth
Secondary phloem in pine stem Tanin cells
Dilatation Tissue • As stem (or root) increases in diameter, circumferential growth of the phloem must occur to keep the tissue from tearing • Two types of dilatation tissue • Proliferative tissue: when axial parenchyma begin to divide and expand • Expansion tissue: when ray parenchyma begin to divide and expand producing a dilated ray
Dilated rays in Tilia show extreme amounts of dilatation tissue