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Plants II. THE SEED PLANTS I. The Vascular Plants. Hypericum sp. Photo by Alan Mulvey. Gymnosperms. Evolution of the Seed Plants. Heterosporous Seed: (embryo & stored food) Evolution of pollen Megagametophyte develops within ovule * Microgametophyte develops within pollen grain *
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Plants II THE SEED PLANTS I The Vascular Plants Hypericum sp. Photo by Alan Mulvey Gymnosperms
Evolution of the Seed Plants • Heterosporous • Seed: (embryo & stored food) • Evolution of pollen • Megagametophyte develops within ovule* • Microgametophyte develops within pollen grain* • Pollen grains evolved by reduction of male gametophyte & protection by pollen walls • Both the male & female gametophytes were retained within the sporophytes
Evolution of the seed • One cell megaspore is retained within megasporangium • It is protected by integument • It develops into mature megagametophyte within the embryo sac • After fertilization, it matures into the embryo (sporophyte)
Generative cell (2 sperms) Tube cell Pollen wall Lilium: Pollen
Generative cell Tube cell Wings Pine: Pollen
Evolutionary transition from a gametophyte to sporophyte dominance From Keeton, 1993
The Gymnosperms: Characteristics • (naked seed)* • Water is not required for fertilization • Mainly pollinated by wind • Reproductive changes increased protection of gametes • Have sunken stoma & thicker cuticles
Two Seeds per Scale Cone Scale Seed Cone
Epidermis Hypodermis Sunken Soma Pine: Leaf
Characteristics Cont.. • Have secondary growth - all trees or shrubs • Mostly evergreen • Xylem has tracheids, lacks vessels (except some Gnetophytes) • Phloem has sieve cells, lacks sieve tube members
Progymnosperms • An extinct group that arose in late Paleozoic • Intermediate between seedless & seed plants • Progymnosperms were ancestors of seed plants • Evolved bifacial V.C. & wood (page 411)
Seed Plants I - Gymnosperms • Conifers – Coniferophyta • Cycads – Cycadophyta • Maidenhair tree – Ginkgophyta • Gnetophytes – Gnetophyta
Conifers: Phylum Coniferophyta • evolved 290 MYO • Constitute the largest group of gymnosperms (50 genera, 550 spp.) • Include the tallest & largest trees ever (Giant redwood) • Conifers (pine, fir, spruce) have great commercial value in temperate zones • Leaves are either • needles, long or short, as in pines, fir, spruce • small flat, scale-like as in junipers, cypresses • Mostly evergreen, monoecious
Pollen Cones Seed Cone White Pine
Turpentine Production Slash Pine: P. elliottii
Giant Red Wood Sequoia http://www.habitas.org.uk/gardenflora/sequoiadendron.htm
Seed Cones Pollen Cones Giant Red Wood http://www.conifers.org/cu/se2/
Leaves Balsam Fir Seed Cone
Larch Developing cones
Common Juniper: Juniperus communis Leaves Tree
White Spruce Pollen cones Young seed cones Pollinated seed cones
Pacific yew: Taxus brevifolia http://www.fw.vt.edu/dendro/dendrology/syllabus/tbrevifolia.htm
Maidenhair tree Phylum Ginkgophyta: • One extant species, Ginkgo biloba (notched leaf)* • Reproduction is dioecious • Ginkgo is resistant to air pollution (city parks) • Pickled seeds are eaten in Asia • Ginkgo is now cultivated, could be in the wild (Asia)
Ginkgo biloba Leaves are broad with dichotomous venation
The Ginkgo tree has properties which make it resistant to nuclear radiation. This tree survived the Hiroshima bombings. www.ginkgo.co.nz/images/ large/ginkgotree.jpg
Male cones Sperms are flagellated, borne in cone-like structure