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변이 (Variation). Nature and Cause of Variation Evolutionary Significance of Variation. 1. Nature and Cause of Variation. Study of Variation within and among individuals, among populations, and among species 자연선택 , 표현형 형질 , 유전자 또는 환경에 따른 변이 정도 , 다음 세대로의 유전 정도.
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변이(Variation) Nature and Cause of Variation Evolutionary Significance of Variation
1. Nature and Cause of Variation • Study of Variation within and among individuals, among populations, and among species • 자연선택, 표현형 형질, 유전자 또는 환경에 따른 변이 정도, 다음 세대로의 유전 정도
Modes of Expression of Variation • Discrete Traits • Quantitative Traits • Threshold Traits • Sexual Dimorphic Traits
Cause of Variation and their Evolutionary Consequences • Variation within individuals • Variation among individuals in random-mating, unstructured populations • Variation in structured populations – genotypes do not interact at random
2. Evolutionary significance of Variation • Quantitative Traits genetic versus environmental causes of phenotype variation
R=h2S • R=h2S • R Response to selection (phenotype change between generation) • H heritability • S Selection differential (before and after selection)
Genetic Variation Additive Genetic Variance Nonadditive Variance - Dominance variance - Cytoplasmic variance • Epigenetic Variance • Interaction of Nonadditive and Additive Genetic Variance
Interconversion of Nonadditive and Additive Genetic Variance • Epistatic variation : Nonadditive Little direct part in response to selection • Epistatic/Alleic variation Loss : Convert to Additive variation • Passing through the bottleneck : Additive Genetic variation increase
Table 2.3 Schematic Example Genotype b loss makes BB only and additive variation
Environmental Variation • Abiotic Environmental Variation • Biotic Environmental Variation • Maternal Environmental Variation
Genotype X Environment Interaction • Cannot be predictated on ths basis of each Genotype and Environment • VP VG + VE ,VP= VGx VE • Diminishing heritability • Constraint evolutionary changes • Reduce potential response to selection