220 likes | 551 Views
변이 (Variation). Nature and Cause of Variation Evolutionary Significance of Variation. 1. Nature and Cause of Variation. Study of Variation within and among individuals, among populations, and among species 자연선택 , 표현형 형질 , 유전자 또는 환경에 따른 변이 정도 , 다음 세대로의 유전 정도.
E N D
변이(Variation) Nature and Cause of Variation Evolutionary Significance of Variation
1. Nature and Cause of Variation • Study of Variation within and among individuals, among populations, and among species • 자연선택, 표현형 형질, 유전자 또는 환경에 따른 변이 정도, 다음 세대로의 유전 정도
Modes of Expression of Variation • Discrete Traits • Quantitative Traits • Threshold Traits • Sexual Dimorphic Traits
Cause of Variation and their Evolutionary Consequences • Variation within individuals • Variation among individuals in random-mating, unstructured populations • Variation in structured populations – genotypes do not interact at random
2. Evolutionary significance of Variation • Quantitative Traits genetic versus environmental causes of phenotype variation
R=h2S • R=h2S • R Response to selection (phenotype change between generation) • H heritability • S Selection differential (before and after selection)
Genetic Variation Additive Genetic Variance Nonadditive Variance - Dominance variance - Cytoplasmic variance • Epigenetic Variance • Interaction of Nonadditive and Additive Genetic Variance
Interconversion of Nonadditive and Additive Genetic Variance • Epistatic variation : Nonadditive Little direct part in response to selection • Epistatic/Alleic variation Loss : Convert to Additive variation • Passing through the bottleneck : Additive Genetic variation increase
Table 2.3 Schematic Example Genotype b loss makes BB only and additive variation
Environmental Variation • Abiotic Environmental Variation • Biotic Environmental Variation • Maternal Environmental Variation
Genotype X Environment Interaction • Cannot be predictated on ths basis of each Genotype and Environment • VP VG + VE ,VP= VGx VE • Diminishing heritability • Constraint evolutionary changes • Reduce potential response to selection