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Roots RAM protected by root cap Roots lack nodes Lateral roots develop deep within root Structural support is less important Endodermis Water absorbing root hairs Cuticle is very thin—if present. Root vs Stem Development. Roots RAM protected by root cap Roots lack nodes
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Roots RAM protected by root cap Roots lack nodes Lateral roots develop deep within root Structural support is less important Endodermis Water absorbing root hairs Cuticle is very thin—if present Root vs Stem Development
Roots RAM protected by root cap Roots lack nodes Lateral roots develop deep within root Structural support is less important Endodermis Water absorbing root hairs Cuticle is very thin—if present Root vs Stem Development
Roots RAM protected by root cap Roots lack nodes Lateral roots develop deep within root Structural support is less important Endodermis Water absorbing root hairs Cuticle is very thin—if present Stems SAM protected by leaf primordia Stems have nodes Leaves & branches develop from surface layer at axillary/lateral bud Structural support is very important No endodermis Cuticle present to prevent dessication Ventilation Root vs Stem Development
What are functions of a stem? • Photosynthesis • Produce leaves • Provide support • Transport food • Transport water • Protection • Elevate leaves
Stems & Leaves • Apical meristem produces more stem, leaves, and flowers • Photosynthesis considerations • Ventilation • Maximize light absorption • Transport photosynthesis products & water • Development of lateral branches • Allow for secondary growth • Develop a new meristem • (topic next week)
Terminal Buds of Wood Plant Contain Shoot Apical Meristem • Scales are modified leaves that protect SAM and leaf primordia from winter, dryness.
Dicot Flowering PlantsAdditions Carrots, fennel, anise, parsley, celery Beans, peas, clover, alfalfa Peppermint, catnip, oregano, thyme, lavender Maples
Sunflower Stem Cross Section “Dicot” Vascular bundles in a ring Pith Distinct collenchyma
Epidermis Collenchyma Cortex Phloem Xylem Pith—parenchyma cells
Epidermis Collenchyma Cortex Phloem Xylem Pith—parenchyma cells
Collenchyma provides support in elongating stem—thickened primary cell walls How do we determine if this is a primary or secondary cell wall?
Formation of primary xylem with some secondary cell walls —cells are elongating as secondary cell walls are produced
Collenchyma—living cells of outer cortex with angular thickening in 1°CW Primary Xylem Water transport & support Inside of phloem Vessel elements, fibers, tracheids, parenchyma Primary Phloem Sucrose & some plant hormone transport Companion cells & Sieve tube members Epidermis Trichomes—protection Reduce water loss Cuticle Waxy layer Reduce water loss Antimicrobial properties Cortex—parenchyma cells Storage Photosynthetic Air spaces Pith—parenchyma storage Primary Stem Tissues
Modified Stems Thorns Honey Locust-Gleditsia Hawthorn-Crataegus
Modified Stems Tendrils Grape-Vitis Cucumber--Cucumis
Monocot Stems have Vascular Bundles that are Scattered Next Slide
New Monocots that are Important Banana, Ginger, Palm, Rice, Wheat, Onion & Garlic
Tulip Pineapple Banana Ginger Orchid Iris
Stem Adaptations to Xeric Environments http://www.usparksonline.com/arizona-national-parks.htm
The Cactus-Like Stem Design“succulents” • Large volume to surface area • Minimize exposure & evaporation • Columnar Shape • Avoid direct sunlight during heat of midday
The Cactus-Like Stem Design“Succulents” • Large volume to surface area • Minimize exposure & evaporation • Columnar Shape • Avoid direct sunlight during heat of midday • Accordion folds • Allow for stem expansion when water is plentiful • Leafless • Reduce surface area and water loss
Desert Lily Liliaceae Desert Milkweed Apocynaceae Desert Grape Vitaceae Desert Poinsettia Euphorbiaceae A True Cactus Cactaceae
Convergent Evolution *unrelated organisms evolve a similar strategy in response to a common selective pressure Desert Lily Liliaceae Desert Milkweed Apocynaceae Desert Milkweed Desert Grape Vitaceae Desert Poinsettia Euphorbiaceae A True Cactus Cactaceae
Cacti have spines in clusters called areoles Spines inflict pain and are difficult to remove.
Stem Review • Shoot apical meristem • Function • Compare with RAM • Tissues of primary stems • Structure & function • Monocot versus Dicot • Convergence evolution of desert Succulents