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Animal Respiration. Lecture #4. Respriation. Involves the diffusion of gases Oxidative respiration consumes O 2 and produces CO 2 Respiration is the process of obtaining O 2 and getting rid of CO 2 This is done by passive diffusion. CO 2 CO 2 O 2 O 2. Composition of Air.
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Animal Respiration Lecture #4
Respriation • Involves the diffusion of gases • Oxidative respiration consumes O2 and produces CO2 • Respiration is the process of obtaining O2 and getting rid of CO2 • This is done by passive diffusion. • CO2CO2 • O2O2
Composition of Air • All O2 in the air is a result of photosynthesis. • O2 producing organisms first appeared in the ocean. • O2 dissolved in water then diffused into atmosphere. • Then, terrestrial plants added O2 to the atmosphere
Composition of Air (II) • Nitrogen (N2)………………78.09% • Oxygen (O2 )………………20.95% • Argon and inert gases………0.93% • Carbon Dioxide (CO2)……..0.03% • Because of convection currents, composition of air is the same throughout the world.
BUT • Composition is the same in a horizontal plane. • Amount of air in a vertical plane is not the same. • As altitude increases, the amount of air decreases.
Air is comprised of molecules. • Therefore, air has • A weight • A pressure • Consider a column or tube of air going from sea level up until there was no more air. • The weight would cause a pressure = a column of 760mm of Hg (mercury) = 1 atmosphere • Weather Channel watchers = 29.92 in
1 Atmosphere = 760mm pressure • Each gas in atmosphere contributes a partial pressure to the total atmospheric pressure. • PN = 78.08% X 760mm = 600.6mm Hg • PO = 20.95% X 760mm = 159.2mm Hg • At sea level, air pressure is 760mm Hg. • But as altitude increases, column gets shorter = air pressure • Partial pressure of O2 (PO) also
Physics of Respiration • Diffusion of O2 across a cell membrane is • Passive • Depends upon the concentration gradient • Limited by distance • (don’t write this down) . We can put this all into a physical equation.
Fick’s Law of Diffusion • Approximates rate of diffusion (R) between 2 regions R = D x A x p/d • D = diffusion constant • A = area over which diffusion takes place • p = concentration gradient [in] vs. [out] • d = distance across which diffusion has to take place
Evolution of Respiration • Single Cells • Limited by size….why? • d = distance of diffusion • Surface [O2 ] would be depleted.
Creating a Water Current • Primitive phyla got rid of O2 depleted water…..how? 1. flagella
2. Movement • In these organisms, ext. [O2 ] never
Increasing Diffusion Surface Area • Advanced invertebrates and vertebrates develop organs that do this. • They R by A • They also d
Fish – Most Efficient • Why? • A (gills) • p (countercurrent flowof water over gills)
Structure of Gills • Water comes in mouth and passes over gills = p
Gill Structure A • Gills segmented into gill filaments = A
Countercurrent Flow • Blood flow in gills opposite that to water flow. • Least [O2 ] blood meets least [O2 ] water at back of gill • Most [O2 ] blood meets most [O2 ] water at front of gill • Result – most efficient blood oxygenation known in nature
Water to Land • More O2 in air than in water • Water = 0.5 – 1% • Land = ~ 21% • Many members of aquatic group (molluscs, crustaceans and fish) came on land. • Two problems to overcome • Gills collapsed….no support • Water evaporation of gills
Two types of respiration on land A. Insects developed trachea.
Insects • Trachea are air-filled passageways connected to all parts of the body. • They close when CO2 levels • water evaporation
Vertebrates • B. Lungs
Amphibians • Earliest terrestrial vertebrates • First to abandon one way respiration • Not a very efficient lung • Could breath with its mouth closed • Could breath through its skin (cutaneous) • Had to have moist skin
Reptiles • More active than amphibians • Require more O2 • Total terrestrial life….no O2 exchange through skin • Dry skin….no dessication (water loss) • Skin is water tight • Lung has A vs. to amphibian
Mammals • O2 consumption much greater than most animals • A • 300 x 106 small sacs called alveoli • Structure of human lung
Lung Pathologiesand Fick’s Law • Emphysema – shortness of breath and inability to do physical exercise (A) • Cystic fibrosis – secretion of thick mucus which D • Respiratory distress syndrome – leading cause of death in premature infants (d)