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http://www.tolweb.org/Arthropoda/2469. mandibulata. Phylum Arthropoda. Subphylum Trilobita Common 500 mya (4000 spp) but extinct by 225 mya Each body segment had a biramous appendage (inner for walking) 3 body tagma Compound eyes with ommatidia. Trilobites flourished for 270 million years.
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http://www.tolweb.org/Arthropoda/2469 mandibulata
Phylum Arthropoda Subphylum Trilobita • Common 500 mya (4000 spp) but extinct by 225 mya • Each body segment had a biramous appendage (inner for walking) • 3 body tagma • Compound eyes with ommatidia
Trilobites flourished for 270 million years. They died.
Subphylum Chelicerata • Include spiders and mites and daddy LL, scorpions, horseshoe crabs • Lack antennae • 1st pair of appendages on the prosoma = the chelicerae (adapted for feeding)
Class Xiphosura: Horseshoe crabs • 4 spp, all marine • Not true crabs (i.e., not crustaceans) • head + thorax = prosoma or cephalothorax • appendages of opisthosoma flattened to become “book gills” for gas exchange Telson
Class Arachnida • Terrestrial • Orders: spiders, mites, ticks, scorpions, ddll • 4 pairs of walking legs, one pair of pedipalps • Head and thorax fused = prosoma • In ticks the prosoma and opisthosoma fuse = carapace
Order Acari: The ticks Lyme’s Disease • deer tick = Ixodes scapularis is a vector for the spirochete, Borrelia burgdorferi, from white-footed mice (reservoir host) and people
Order Scorpiones • Most ancient arachnid (and therefore most primitive terrestrial arthropod) • Also the largest arachnid is a scorpion (18cm)
Class Arachnida, Order Scorpiones A delicacy in Bankok, Thailand
Opisthosoma Mesosoma Prosoma Metasoma Eyes Chelicera Aculeus Telson Chela Pedipalp
Class Arachnida:Order Aranea (Spiders) • Book gills internalized: = book lungs • Spiracles and trachea • Gas exchange does not require blood • Spiders are specialized predators • Poison glands, hollow chelicerae • Silk glands, web building • Spinneret are modified appendages • 7 separate silk glands make different kinds of silk
book lung
O. Pseudoscorpiones No stinger Dispersal by phoresy (hitchhike on houseflies) Poison in chelae for immobilizing prey
Order Amblipygi • First legs held out while they scurry sideways • serve as tactile and chemoreceptors Tailless whip scorpions are common in Costa Rica
Subphylum Mandibulata • Class Myriapoda • Order Chilopoda • Order Diplopoda • Class Insecta (Hexapoda) • 33 orders • Class Crustacea • 6 subclasses
Myriapoda • 1st head segment bears antennae • 2nd head segment vestigial (no antennae) • In crustacea this segment bears 2nd pair of antennae • Gas exchange by trachea, spiracles on each body segment • Not closable, risk of water loss • Cuticle not waterproof – more water loss • Excretion by malpighian tubules • Repugnatorial glands on ventrum
Claws (modified legs) with poison glands for immobilizing prey • Uniramous appendages, one per segment • Spiracles cannot be closed to control water loss mandibles cephalon Non-locomotory anal legs Defensive Sensory Aggressive
Diplopoda • Diplosegments give the appearance of 2 pairs of legs per segment • Lack poison claws • Some produce cyanide defensive secretions