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The Visual System: Color Vision. Lecture 16. The Trichromatic Theory. Young-Helmholtz (1802) 3 types of color receptors Cones Differential sensitivity to light wavelengths red (long) green (medium) blue (short) ~. The Trichromatic Theory. Perceived color
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The Visual System:Color Vision Lecture 16
The Trichromatic Theory • Young-Helmholtz (1802) • 3 types of color receptors • Cones • Differential sensitivity to light wavelengths • red (long) • green (medium) • blue (short) ~
The Trichromatic Theory • Perceived color • overall pattern of stimulation • Like mixing paint • Negative After-image?
The Trichromatic Theory? • What colors do you see? • How can you see colors that weren’t there? • Negative After-image • Does it fit Trichromatic Theory? ~
Hering Opponent Process Theory • Competing theory • Center-Surround organization • BP, RGC, & LGN • Antagonistic for color • Red-Green • Blue-Yellow • Black-White
Hering Opponent Process Theory • Center- surround antagonistic • Each color can be excitatory (+) or inhibitory (-) • 12 combinations total ~
Which Theory? • Both are correct in retina • Photoreceptors: trichromatic • Higher levels: Opponent Process • BP, RGC, & LGN • More complex at cortical level • Retinex Theory ~
Retinex Theory • Perception of visual stimulus • subjective • includes context & past experiences • color & brightness constancy • Color constancy • e.g., grass in sun vs. grass in shade • perceived as same color green • e.g., Rubic’s cube ~
Lateral Geniculate Nucleus • 6 layers • dorsal ventral • 6 1 • RFs center-surround • Input from each eye • monocular • 2, 3, 5 from the ipsilateral eye • parallel processing ~
LGN: Parallel Processing • Parvocellular system • layers 3-6 • small RFs • info from cones • color & form • Magnocellular system • Large RFs • layers 1 & 2 • info from rods • form only, no color ~
V1 Organization • 6 layers • Most input Layer 4 • 1st binocular receptive fields • cells get input from both eyes • ocular dominance • Modular organization • Blob cells: color processing • Wavelength specific ~
Modular V1 Organization L R L B L O B 2 & 3 4 LGN 5 & 6
Blob cells & Color Perception • Same color perceived differently • Double Opponent Process Cells • Different from RGC & LGN • center-surround • Center: • red excitation • green inhibition • Surround: • red inhibition • green excitation ~
Double-Opponent Process Blob Cells R-G+ R-G+ R+G- R+G-
Higher Level Color Processing • V1 V2 V4 • V4 in medial occipital cortex • V4 damage • disrupts color constancy • achromatopsia • unilateral – 1 visual field • bilateral – no color perception ~