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Chapter 46: Community Structure and Biodiversity. Community. All the populations that live together in a habitat Type of habitat shapes a community’s structure. Factors Shaping Community Structure. Climate and topography Available foods and resources Adaptations of species in community
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Community • All the populations that live together in a habitat • Type of habitat shapes a community’s structure
Factors Shaping Community Structure • Climate and topography • Available foods and resources • Adaptations of species in community • Species interactions • Arrival and disappearance of species • Physical disturbances
Niche Sum of activities and relationships in which a species engages to secure and use resources necessary for survival and reproduction
Realized & Fundamental Niches • Fundamental niche • Theoretical niche occupied in the absence of any competing species • Realized niche • Niche a species actually occupies • Realized niche is some fraction of the fundamental niche
Species Interactions • Most interactions are neutral; have no effect on either species • Commensalism helps one species and has no effect on the other • Mutualism helps both species
Species Interactions • Interspecific competition has a negative effect on both species • Predation and parasitism both benefit one species at a cost to another
Symbiosis • Living together for at least some part of the life cycle • Commensalism, mutualism, and parasitism are forms of symbiosis
Mutualism • Both species benefit • Some are obligatory; partners depend upon each other • Yucca plants and yucca moth • Mycorrhizal fungi and plants
Yucca and Yucca Moth • Example of an obligatory mutualism • Each species of yucca is pollinated only by one species of moth • Moth larvae can grow only in that one species of yucca
Mycorrhizae • Obligatory mutualism between fungus and plant root • Fungus supplies mineral ions to root • Root supplies sugars to fungus
Sea Anemone and Fish Fig. 46-4, p.823
Competition • Interspecific - between species • Intraspecific - between members of the same species • Intraspecific competition is most intense
Forms of Competition • Competitors may have equal access to a resource; compete to exploit resource more effectively • One competitor may be able to control access to a resource, to exclude others
Interference Competition Least chipmunk is excluded from piñon pine habitat by the competitive behavior of yellow pine chipmunks Least Chipmunk Yellow Pine Chipmunk
Competitive Exclusion Principle When two species compete for identical resources, one will be more successful and will eventually eliminate the other
Gause’s Experiment Paramecium caudatum Species grown together Paramecium aurelia Figure 47.6 Page 825
Hairston’s Experiment • Two salamanders species overlap in parts of their ranges • Removed one species or the other in test plots • Control plots unaltered • 5 years later, salamander populations were growing in test plot
Resource Partitioning • Apparent competitors may have slightly different niches • May use resources in a different way or time • Minimizes competition and allows coexistence Figure 47.8 Page 825
bristly foxtail Indian mallow smartweed Fig. 46-8b, p.825
Predation • Predators are animals that feed on other living organisms • Predators are free-living; they do not take up residence on their prey
Coevolution • Joint evolution of two or more species that exert selection pressure on each other as an outcome of close ecological interaction • As snail shells have thickened, claws of snail-eating crabs have become more massive
Predator-Prey Models • Type I model: Each individual predator will consume a constant number of prey individuals over time • Type II model: Consumption of prey by each predator increases, but not as fast as increases in prey density • Type III model: Predator response is lowest when prey density is lowest
Canadian Lynx and Snowshoe Hare • Show cyclic oscillations • Krebs studied populations for ten years • Fencing plots delayed cyclic declines but didn’t eliminate them • Aerial predators, plant abundance also involved • Three-level model
Prey Defenses • Camouflage • Warning coloration • Mimicry • Moment-of-truth defenses
Camouflage Fig. 46-11a, p.828
Camouflage Fig. 46-11b, p.828
Camouflage Fig. 46-11c, p.828
Mimicry Fig. 46-12a, p.829
Mimicry Fig. 46-12b, p.829
Predator Responses • Any adaptation that protects prey may select for predators that can overcome that adaptation • Prey adaptations include stealth, camouflage, and ways to avoid chemical repellents
Parasitism • Parasites drain nutrients from their hosts and live on or in their bodies • Natural selection favors parasites that do not kill their host too quickly
Kinds of Parasites • Microparasites • Macroparasites • Social parasites • Parasitoids