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Asterids: Asteraceae

Asterids: Asteraceae. Asterids. Very large, diverse group 10 orders, many families Apomorphies: iridoid compounds sympetalous corolla ovules: one integument and the megasporangium is generally one cell thick. Asteraceae. Probably the most evolved of the families Two types of flowers

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Asterids: Asteraceae

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  1. Asterids:Asteraceae

  2. Asterids • Very large, diverse group • 10 orders, many families • Apomorphies: • iridoid compounds • sympetalous corolla • ovules: one integument and the megasporangium is generally one cell thick

  3. Asteraceae • Probably the most evolved of the families • Two types of flowers • Ray • Disk • Herbs in our area, but can be trees in tropics • Very diverse: 19,000 species world-wide • Many flowers on the head inflorescence • Old name is compositae – b/c a composite of flowers

  4. Asteraceae • Inflorescence a head (capitulum): (usu.) many flowers arising from a compound receptacle, subtended by inflorescence bracts: involucral bracts or phyllaries, collectively termed the involucre. • Calyx modified as pappus. • Fruit an achene.

  5. Floral Formula

  6. Asteraceae – World-wide

  7. Typical Asteraceae Flower

  8. Cross-section of Asteraceae Flower

  9. More detail on disk flower

  10. Note the chaff – bracts below each individual flower

  11. Fruits are achenes

  12. Sunflower “seeds” = achenes

  13. Echinacea sp. – praire cone flower

  14. Liatris flowers

  15. Close-up of achene fruit with pappus

  16. Taraxacum officinale

  17. Close-up of Taraxacum head

  18. Taraxacum fruits

  19. Close-up of Taraxacum flower

  20. Five types of heads: 1) discoid, with only disk flowers; 2) disciform, with central disk flowers and marginal, eligulate female flowers; 3) radiate, with central (bisexual or male) disk flowers and peripheral (female or sterile) ray flowers; 4) ligulate, with all ray flowers (typically with 5-toothed corolla apices); 5) bilabiate, with all bilabiate flowers.

  21. Bilabiate flower posterior lip anterior lip Acourtia microcephala Trixis californica

  22. Ray flowers (heads ligulate = all rays) ligulate corolla Rafinesquia neomexicana Malacothrix californica

  23. Disk flower: heads discoid disk corolla Palafoxia arida Chaenactis gabriuscula Psathyrotes ramosissima

  24. Disk flowers: heads disciform(2 types of disk fls., same or different heads) male heads female heads Ambrosia chamissonis

  25. Heads radiate: inner disk & outer ray fls. disk flowers ray flowers Xylorhiza orcuttii Encelia farinosa

  26. Some heads are "chaffy" ovary disk flower Encelia californica chaff : bracts subtending flowers

  27. Involucre morphology two whorls many whorls one whorl Senecio vulgaris Coreopsis maritima Encelia californica

  28. Involucre morphology Phyllaries spiny & squarrose Phyllaries spiny Circium vulgare Silybum marianum

  29. Pappus: modified calyx

  30. Pappus: modified calyx beak capillary bristles, borne atop "beak" capillary bristles: barbellate capillary bristles: plumose

  31. Asteraceae • Because the family is so large, it is divided into two Subfamilies which are then divided into tribes: 12 to 17 in the whole family • Aster = Subfamily Asteroideae • Six tribes we’ll see • Dandelion = Subfamily Cichorioideae • One tribe

  32. Tribes of the Aster Subfamily • Heliantheae • Asterae • Anthemideae • Senecioneae • Gnaphalieae • Eupatorieae • Cynareae

  33. Tribe Heliantheae – Sunflower tribe • Opposite leaves • Very resinous • Bracts attached to individual flowers (chaffy receptacle) • Both types of flowers • Sunflower (Helianthus) and Echinacea

  34. Heliantheae

  35. Tribe Asterae – Aster Tribe • No bracts at the bottom of individual flowers • Example is genus Aster, generally fewer than 20 “petals” or ligulate flowers • Similar to above tribe, but without resinChrysothamnus (Rabbitbrush or chico) • Another important genus is Erigeron aka fleabane daisy

  36. Chyrsothamnus sp. rabbitbrush

  37. Erigeron elatior

  38. Tribe Anthemideae – Chamomile Tribe • Multiple layers of phyllaries of different lengths • Richly aromatic • Thin, dry, and translucent (i.e. scarious) bracts surrounding the flower base • Artemisia (sagebrush is in this tribe) • Achillea, Chrysanthemum

  39. Involucre morphology two whorls many whorls one whorl Senecio vulgaris Coreopsis maritima Encelia californica

  40. Chrysanthemum

  41. Achillea millefolium - yarrow

  42. Tribe Senecioneae – Senecio or Groundsel Tribe • Silky pappus around each flower • Pappus is usually pure white and very abundant • Usually naked receptacle • Senecio

  43. Senecio triangularis

  44. Tribe Gnaphalieae – Everlasting Tribe • Also has scarious bracts around flower base • No ligulate flowers, only tubular flowers • Example is cudweed or everlasting (Gnaphalium)

  45. Pussytoes in Gnaphalieae

  46. Tribe Eupatorieae – Boneset Tribe • “Baseball bat stigmas” • No ligulate flowers • Usually naked receptacle • Snakeweed, Gutierrezia • Liatris, common ornamental

  47. Gutierreziasarothrae – broom snakeweed

  48. Liatrispunctata

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