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Earliest Hominins. Hominin Phylogeny. Genus Homo. Archaic. Primitive. Mosaic Evolution. Modern human characteristics Appeared at different times Evolved at different rates Extinct hominids show a mosaic of derived and ancestral characteristics. Hominin Evolutionary Sequences.
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Hominin Phylogeny Genus Homo Archaic Primitive
Mosaic Evolution • Modern human characteristics • Appeared at different times • Evolved at different rates • Extinct hominids show a mosaic of derived and ancestral characteristics
Hominin Evolutionary Sequences • Locomotion • Shift from inefficient to more efficient bipedality ca. 2 mya • Dentition • Enamel thickening ca. 4 mya • Gradual reduction of anterior dentition 4.5 mya to present • Gradual reduction of posterior dentition ca. 2.5 mya to present • Encephalization • Limited encephalization from 4 mya to 2.5 mya • Marked encephalization from 2.5 mya to ca. 150 kya
Skeletal Indicators of Bipedality • Pelvis • Distribution of body mass • Relative limb-length • Position of foramen magnum • Lumbar vertebrae • Foot
Sahelanthropustchadensis • Age – estimated to be between 6 and 7 mya • Fossil remains • Nearly complete cranium • Several fragmentary mandibles • Isolated teeth • Cranial capacity – 320-380 cc • Classification uncertain – several possibilities • Position of foramen magnum suggests non-bipedal
Sahelanthropus tchadensis • Chimp-sized brain • Hominin-like brow ridges • Canine wear pattern not ape-like • Thick hominin-like mandible
Orrorin tugenensis • Age ca. 6mya • Fossils remains – mandibles, femur, humerus, large molars, small canines • Femur suggests bipedality • Humerus suggests tree-climbing skills (but not brachiation) • Long bones suggest chimpanzee size • Teeth resemble later hominids
GRACILE & ROBUST AUSTRALOPITHECINES “Robust” (aethiopicus) “Gracile” (afarensis)
Australopithecus africanus • STS 5
“gracile” vs “robust” archaic hominins • Au. africanus • P. boisei
Australopithecus ghari • Age ca. 2.5 mya • Later Australopithecine • Limbs suggest bipedality • Incipient megadontia • Found with stone tools and butchered animal remains