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Basal Tricolpates. Finally, true dicots. Basal Angiosperms, view 1. Basal Angiosperms, view 2. Characters of Tricolpates. Tricolpate pollen Floral parts whorled, members alternating Staminal filaments slender Sieve element plastids with starch grains Nuclear & chloroplast sequence data.
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Basal Tricolpates Finally, true dicots
Characters of Tricolpates • Tricolpate pollen • Floral parts whorled, members alternating • Staminal filaments slender • Sieve element plastids with starch grains • Nuclear & chloroplast sequence data
Basal Tricolpates – Utah View • Ranunculales, order of buttercup family, is important • Ranunculaceae, Berberidaceae, Papaveracae • Platanales of some interest • Proteales – fascinating but mostly southern hemisphere
Ranunculales • Mostly herbaceous • Toothed to lobed or compound leaves • Floral parts distinct, calyx, often many stamens, ovary superior, seeds with tiny embryos and much endosperm • Alkaloid berberine • Nuclear and chloroplast DNA
Papaveraceae (inc. Fumariaceae) • Deciduous sepals • Carpels 2, connate • Fruit a capsule • Nearly all have laticifers • Also has properties of Ranunculales
Papaveraceae (2) • Herbs or shrubs • Laticifers present (opium), alkaloids • Lvs simple but lobed or dissected, pinnate venation • Flowers bisex, radial or bilat, Sepals 2, deciduous, petals 4 or 6 (often crumpled), sometimes united distally, stamens many to 2*3, carpels 2, united, superior ovary, placentation parietal, fruit a capsule
Dendromecon Dicentra Corydalis Lump or split? Argemone
A family of exceptions Mostly herbaceous, some shrubs, vines Lvs usually alt or spiral, usually simple, oft lobed or dissected, usually pinnate venation, usually no stipules Flowers usually bisex, usually radial, usually not 3-merous, tepals distinct, calyx usually distinct; stamens usually many, carpels usually 5+, US distinct, ovary superior Fruit follicles (dehiscent, >1 seed) or achenes (indehiscent, 1 seed) Ranunculaceae
Many stamens No hypanthium No stipules Ranunculaceae
Berberidaceae • Herbs or shrubs (ours), yellow wood (berberine) • Lv usually alt to spiral, simple to compound, entire to serrate or spinose-serrate • Flrs bisex, rad, usually 3 merous, calyx distinct, inner petals (staminodes?) nectar-producing, stamens 4-many,anthers opening by flaps, carpel 1, ovary superior; fruit usually berry
Platanaceae • Trees, monoecious • Branched hairs • Lvs alt or spir, usually palmate, gland tooth, petiole covers axil bud, stipules • Flr unisex, reduced. Sep 3-7,pet 3-7, st 3-7, carp 5-9, distinct, ov sup. Fruit achene, with bristles
web1.p15132387.pureserver.info/.../ Platane/ www.hope.edu/.../BasalEudicots/ Platanaceae Platanaceae www.hope.edu/.../BasalEudicots/ Platanaceae