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Bivalves and Cephalopods. About one thousand species live in fresh water. Zebra Mussel Zebras on native clams (Unionidae). Best understood in the context of their largely sedentary adult life style. umbo. ctenidium. Adductor muscle. Right valve. Labial palps.
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About one thousand species live in fresh water Zebra Mussel Zebras on native clams (Unionidae)
Best understood in the context of their largely sedentary adult life style
umbo ctenidium Adductor muscle Right valve Labial palps Excurrent Incurrent siphons mouth Foot Mantle edge
Two principal types, distinguished by their Ctenidia and mode of feeding Class Bivalvia Protobranchia Llamellibranchia Filibranchia Eullamellibranchia
Ctenidia of Bivalves are modified for feeding except in one group: the Protobranchs
In most bivalves ctenidia are sheet-like with high S.A., and are used for feeding
Captured particles are moved to a ciliated ventral groove and then as a mucus strand to the mouth. http://www.biology.ualberta.ca/facilities/multimedia/index.php?Page=252
Summary of Water Flow and Feeding http://web.uconn.edu/jevanward/video/musselsum.MPG The sheetlike, arrangement of ctenidia in most bivalves is known as the Llamellibranch condition
There are two basic type of Llamellibranch ctenidia in bivalves Filibranch (in mussels) Eullamellibranch (most clams)
Living Cephalopods Squid Nautilus Cuttlefish Octopods --The class as a whole is adapted for swimming, suspended in the water by buoyancy mechanisms. -- allare carnivores, occupying the same niche as fishes
Architeuthys Giant squid up to 20 m long Mesonycheteuthis Colossal squid up to 14 m major prey item of sperm whales Dosidicus Humboldt squid, smaller but numerous (2 m) and comes to surface to feed
Ammonites Largest is 3 m in diameter
Sub Class Nautiloidea 5-6 species remain from diversity in geologic past Live in the last chamber of a multi-chambered shell Move by forcing water through a siphon.
Sub Class Nautiloidea How does a Nautilus adjust and maintain its buoyancy in water?
Cuttles, Squids and Octopuses • tendency is reduction of shell. • compensated for by selective ionic control; accumulate ammonium • other adaptations to shell loss and life style
cuttle chameleons of the sea most specialized, agile and cryptic octopus
Specifically, compensation takes the following forms: Stealth and other defenses Acute sensory structures Speed Cunning
A. Stealth and other defenses Ink Sack : ink with melanin and mucus also dopamine, L-DOPA, tyrosinase Pseudomorphs
A. Stealth and other defenses Chromatophores: each cell contains a different pigment which may be red, orange, brown, black, yellow or blue
Cephalopod Chromatophore Black, brown, orange or yellow colors Also iridiophore reflective cells Most cephalopods are completely colorblind neurophilosophy.files.wordpress.com
B. Acute sensory structures -- Vision -- Lateral line analogs
C. Speed Rival fish in ability to swim, attaining burst speeds of 5-10 m second sustained for short distances at speeds of up to 40 km hr. Burst speed is achieved by jet propulsion More gradual swimming is fin undulation
C. Speed : complex nerve control of locomotion Assures synchronized contraction of mantle muscles
C. Speed : circulatory system Hearts Requires a closed, elaborate circulatory system
D. Cunning -- Relative to body size, the brain is larger than that of most fish
Experiments with Octopus vulgaris show their ability to learn. They learn to choose colors not associated with shock. Observers choose properly colored objects after watching demonstrators