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Somatosensory Systems. cutaneous proprioceptive. Adequate Stimuli. Thermal (infrared radiation, contact). Touch (light touch, pressure, vibration). Pain and Itch (chemical, thermal, mechanical). Proprioception (mechanical; stretch or pressure). Cutaneous subsystems. epicritic
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Somatosensory Systems cutaneous proprioceptive
Adequate Stimuli Thermal (infrared radiation, contact) Touch (light touch, pressure, vibration) Pain and Itch (chemical, thermal, mechanical) Proprioception (mechanical; stretch or pressure)
Cutaneous subsystems epicritic location vibration texture shape protopathic pain temperature itch and tickle
Epricritic, or non-pain Somatosensation
Meissner’s Merkel’s 60 hz vibration Pressure Riffini’s Free nerve ending Pacinian Pain Stretch 200 hz vibration
As in the retina, receptive fields vary in size. Smaller receptive fields = greater acuity two-point discrimination
Center-surround organization of cutaneous receptive fields results in lateral inhibition, as in retina Serves to enhance contrast
Protopathic, or pain Somatosensation
Pain Receptors Called Nociceptors • Free nerve endings that respond to: • mechanical stimuli • thermal stimuli • chemical stimuli, or • all three • (polymodal receptors)
Free nerve endings of unmyelinated C fibers or thinly myelinated Aδ fibers
Cutaneous classified by conduction velocity Proprioceptive classified by axon diameter
SubstanceEffect Potassium activation Bradykinin activation Histamine activation Prostaglandins sensitization Substance P sensitization
Gate control theory of pain control
Parallel Processing in the Somatosensory System Lemniscal System (non-pain; epicritic) Extralemniscal System (pain; protopathic) Spinothalamic pathways Neospinothalamic Paleospinothalamic Spinomesencephalic
Neospinothalamic Paleospinothalamic Spinomesencephalic
Sensory System Summary 1. Sensory systems detect change over space (lateral inhibition to enhance contrast) over time (rapidly adapting) 2. Detect “features” 3. Structures are laminated (cells in layers) 4. Parallel pathways 5. Hierarchical processing 6. Topographical organization 7. Non-uniform receptive fields 8. Extreme sensitivity, wide dynamic range 9. Non-linear response