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Species number dynamics by Stenseth S9. Species do not reproduce itself, supplied by exotic source. ≈ gap dynamics with seed-bank ≈ metapopulation dynamics, If so, we remind global species richness in local species number dynamics. Species-abundance pattern by Tekeshi S10.
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Species number dynamics by Stenseth S9 • Species do not reproduce itself, supplied by exotic source. • ≈ gap dynamics with seed-bank≈ metapopulation dynamics, • If so, we remind global species richness in local species number dynamics.
Species-abundance pattern by Tekeshi S10 • Statistically null model (Random fluctuation) explains well, including fossil record. • What difference in patterns between niche division by specialists or guild formation by generalists?
Island biodiversity and speciation by Chiba S11 • Minority species advantage in predation risk • Sexual selection (female choice) in hermaphrodite land snails • Sequential adaptive and maladaptive radiation
Pop Ecol Symposia have made a big history… • After PES in 1987 at 熱川 & 1989 広島 • Iwasa et al. (1994 Nature) sexual selection model • Taper & Case (1992 Evol.) taxon cycle model • Matsuda et al. (1995 Evol) ES fitness minima • based on a new quantitative genetic model, from Roughgarden (1987), Vincent & Brown (1988)
Lande (1976) Freq-independent Normal phenotype distrib. Iwasa et al. (1991) Freq-dependent Small phenotype variance Two quantitative genetic model
Pomiankowski’s sexy son modelwith mutation bias • X: male sexy trait, y: female choice
Taper & Case(1992) Doebeli S4 Taxon cycle Rummel & Roughgarden (1987) Fitness, population size Size advantage
Adaptive landscape changes with mean trait value●(Evol. “Stable” Fitness Minima)→ driving speciation (Doebeli S4) Abrams, Matsuda & Harada (Evol. Ecol 1992)
Natural selection S1 Gavrilets (spatial structure) S3 Kaneko (synergistic effects) S4 Doebeli (freq. dependent sel. & assortative mating.) Is Speciation driven by natural- or sexual-selction? • Sexual selection: • S2 Takimoto • (≈sexy-son & less sexy hybrid) • E23 Tanaka • (≈good gene & migration)
Hybrid is … • Gavrilets “disadvantage in freq-indep-natural selection (Holey adaptive landscape). AA>Aa<aa • Takimoto “less sexy, even advantage (=less costly) in natural selection AA<Aa>aa
Why do African cichlids have intra-specific variation? • Maintained by prey-predator relationship • Frequency dependent natural selection without assortative mating
This year symposium will open a new page of Population Ecology History. • Thanks to the Organizing Committee, All speakers and audience.