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Ch 23: Understanding Diversity: Systematics. phylogeny systematics taxonomy taxon. How to categorize life? life vs. non-life (?prehistory) plants (non-motile) vs. animals (motile) Greek philosophers including Aristotle ( born 384 - died 322 BC ).
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phylogeny systematics taxonomy taxon
How to categorize life? life vs. non-life (?prehistory) plants (non-motile) vs. animals (motile) Greek philosophers including Aristotle (born 384 - died 322 BC)
add Protista/Protoctista & sub-group Monera (Haeckel 1866) prokaryotic vs. eukaryotic cells (Chatton 1937)
reorganized classification into 4 kingdoms (Copeland 1930s?) 5 kingdoms; adding Fungi (Whittaker 1959)
3 domains (Carl Woese 1977) Carol J. Bult
further elaborated by Mitchell Sogin (early eukaryotes) & many others
-> 6 kingdoms (compromise) still argued: Lake’s 4 domains, Cavalier-Smith & others’ > 6 kingdoms
horizontal gene transfers genome fusions clades?
Groups of organisms on a phylogenetic tree (excluded)
Homologous structures 18-14 modified leaves 18-13 terrestrial vertebrate forelimbs
Homoplasies = analogous structures Australian marsupial “mole” 18-16 spiky plant parts N American placental mole
Phenetics/evolutionary systematics numerical taxonomy eg: Birds -> different enough to be a separate class from reptiles
Cladistic systematics clade cladogram synapomorphy vs. (sym)plesiomorphy outgroup parsimony eg: Birds -> sub-group of Reptile class
Morphology: Developmental data eg: early animal embryo cleavage (30-5)
Biochemistry: Lipids 2 species of Pseudomonas
Biochemistry: Proteins eg: Isozyme patterns from a fungus
Biochemistry: Proteins eg: amino acid sequence comparison
Biochemistry: Proteins eg: 3D structure comparison myoglobin
Biochemistry: DNA Sequence analysis Relationships between canids using DNA sequences
Stoneking, et al. Mitochondrial DNA study of humans
Other kinds of trees Phylogram - shows genetic change Ultrametric tree - shows time
Methods of generating trees Distance Parsimony Maximum likelihood