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9. Almost Rosids to Rosids. Santalales. Lots of parasitism Polyacetylenes 1-seeded ovules, pendulous and free-central placentation Molecules In ours, flowers not prominent Families not on required list. “Santalaceae”. Hemiparasites Root parasites. Viscaceae. Epiphytic parasites
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Santalales • Lots of parasitism • Polyacetylenes • 1-seeded ovules, pendulous and free-central placentation • Molecules • In ours, flowers not prominent • Families not on required list
“Santalaceae” • Hemiparasites • Root parasites
Viscaceae • Epiphytic parasites • Haustoria • Stems jointed • Lvs opp, oft. Coriaceous or succ. • Flrs unisex, inconspic, radial • T3-4A3 (no fil),G3-4, inf. Viscous berry • Passive gametophyte • Pantrop + some temperate
Loranthaceae • Us. Epiphytes • Lv opp or almost so • Flrs bisex, rad or bilat • Ca rimCo(3)5-6(9), distinct or conn, oft. Red or yellow, A=Co, opp, fil adnate • Aggr. gametophyte • Tropical
ROSIDS • Weakly supported group • Also some lack of clarity of relationships within • Lots of representatives in our part of the world • Hypanthium features in many families • Radial symmetry common
Saxifragales • Some families chiefly (only?) on molecular data • Hypanthium, 2-united carpels, glandular leaf teeth • Retained: 5-merous flowers with separate parts
Saxifragaceae • Two interpretations Shaw – as in text; Welsh – much broader (older) • Herbs, often basal leaves, alt. NO STIPULES • Flr rad to bilat, HYPANTH, • Ca4-5 sometimes connate, Co4-5, oft. Clawed, St 3-10. G2(-5), connate, varying position, axile, oft. nectar disk • Fr. Capsule or follicle
Geraniales • Five families, 1 in north Utah • Morphological features of order • Leaf margins with glandular teeth • Inflorescence cymose • Calyx persistent in fruit • A few other characters • Anomocytic stomates • Vessel elements with simp[le perforations
Geraniaceae • Remarkable feature is fruit • Herbs to subshrubs; stems usually jointed at nodes • Hairs glandular, with aromatic oils (I.e., smelly plants) • Stipules present, leaves with gland. Teeth • K5C5A(5)10,15, filaments often slightly connateG5conn • Fruit a schizocarp, persistent style column
Persistent style Note the fruits in picture above and on the right
Malpighiales • This order contains 7.8% of the diversity of angiosperm diversity • There does not seem a whole lot tying it together, see: http://www.mobot.org/MOBOT/Research/APweb/welcome.html The Angiosperm Phylogeny Group’s home page