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Control Systems in Plants. Plant Hormones. Coordinates growth Coordinates development Coordinates responses to environmental stimuli. Plant Hormones. Auxin (IAA) Cytokinins Gibberllins Abscisic Acid Ethylene Oligogaccharins Brassinosteroids. Auxins. Stimulates stem elongation
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Plant Hormones • Coordinates growth • Coordinates development • Coordinates responses to environmental stimuli
Plant Hormones • Auxin (IAA) • Cytokinins • Gibberllins • Abscisic Acid • Ethylene • Oligogaccharins • Brassinosteroids
Auxins • Stimulates stem elongation • Stimulates root growth • Stimulates differentiation and branching • Stimulates development of fruit • Stimulates apical dominance • Stimulates phototropism and gravitropism
Auxin Control • Auxin stimulates growth • Auxin block on right causes cells to elongate and the plant bends left • Auxin block on left causes cells to elongate the the plant bends right
Polar Transport • Auxin is transported from apex to shoot • Cell wall is acidic • auxin ion picks up H+ • diffuses across plasma membrane • Cytosol is neutral • auxon loses H+ • uses energy to pump out H+ to maintain pH • leaves cell through carrier proteins
Acid Growth Hypothesis • Proton pump stimulated by auxin which lower pH of wall • Hydrogen Ion activates Enzyme • Enzyme breaks hydrogen bonds in cellulose • Wall takes up water and elongates
Auxin Others • Promotes secondary growth by encouraging vascular cambium and secondary xylem • Promotes adventitious root at the base of a cut stem • Promotes fruit growth without pollination (seedless tomatoes)
Cytokinins • Stimulates root growth • Stimulates cell division and differentiation (with auxins) • more cytokinin - shoot buds develop • more auxin - roots develop • Stimulates germination • Delays Senescence
Gibberellins • Promotes seed and bud germination • Promotes stem elongation • Promotes leaf growth • Stimulates flowering and fruits • (with auxin)
Abscisic Acid • Inhibits growth (with gibberellins) • Closes stomata under water stress • Permits dormancy
Ethylene • Promotes fruit ripening • Controls Abscission • (with auxin)
Oligosaccharins • Triggers defense responses against pathogens • Regulates growth
Brassinosteroids • Required for normal growth and development
Signal-Transduction Pathways • A hormone binds to receptor and stimulates secondary messengers which Activates cell responses
Plant Movements • Phototropism • Gravitropism • Thigmotropism
Plant Movement • Rapid Leaf Movement • drop in turgor pressure within pulvini (at joints of the leaf) • sent by action potentials • Sleep Movements • cells on opposite sides of pulvinus control the movement
Daily and Seasonal Responses • Circadian Rhythm • Photoperiodism • controls flowering (short-day vs. long-day) • critical night length
Flowering Hormones • Experiment indicates the presence of some type of flowering hormone
Phytochromes • Function as photoreceptors / red (660nm) to far red (730nm) • Activates kinases (regulatory proteins)
Plant Responses to Environmental Stress • Water Deficit • Oxygen Deprivation • Salt Stress • Heat Stress • Cold Stress • Herbivores
Water Deficit • Slows Transpiration • stomata close • release of abscisic acid • Inhibits growth of young leaves • Change leaf shape • Deeper root growth
Oxygen Deprivation • Aerial roots • Production of air tubes
Salt Stress • Salt glands • Compatible solutes
Heat Stress • Heat-shock proteins
Cold Stress • Altering the lipid composition of their membranes • Changes in the solute composition of the cytosol
Responses to Herbivores • Produce Canavanine • Recruitment of predatory animals
Defense Against Pathogens • Gene-for-gene recognition
Defense Against Pathogens • Hypersensitive response (HR) • phytoalexins are released • PR proteins are released • Seal off infected area • Systemic acquired resistance (SAR) • Salicylic acid released due to death of cell • Activates a Signal Transduction Pathway • Activates SAR