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Fall 05 skip. acclimitization. Acclimatization effects on final rectal temperature, heart rate, and sweat rate, for a group of men exposed daily to 2 hr of hard work in a hot environment. (after Leithead CS, Lind AR. Heat Stress and Heat Disorders. 1964. Philadelphia, Pa.: FA Davis Co, 304.
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Fall 05 skip acclimitization Acclimatization effects on final rectal temperature, heart rate, and sweat rate, for a group of men exposed daily to 2 hr of hard work in a hot environment. (after Leithead CS, Lind AR. Heat Stress and Heat Disorders. 1964. Philadelphia, Pa.: FA Davis Co, 304.
EMILI GARCÍA-BERTHOU Ontogenetic Diet Shifts and Interrupted Piscivory in Introduced Largemouth Bass (Micropterus salmoides) Internat. Rev. Hydrobiol. 87 2002 4 353–363 acclimitization Fall 05 skip
acclimitization Fall 05 skip
acclimitization Fall 05 skip
Fall 05 skip acclimitization
Fall 2005 Life Table
Fall 05 Start Notes on Evolution,Natural Selection, and Speciation
One picture of evolution What is going on before a split? How did the splits occur? What is speciation?
Mainland Types of evolution ↑NA migration • Macro • Micro • Gene flow (E and I) • In isolated populations • Migrating indivs. start breeding • Immigrants can add new alleles • Not necessarily random • Genetic Drift • Random mating in small populations • Causes relative success of only a fraction of individuals • Population bottlenecks (large population) • Founder effects (small population) ↓NA
Natural Selection • Differential contribution of offspring to next generation by individuals in a population • Not Random
Recall: Normal curve of phenotypic variation Height of students in class
Another way to look at it… • Directional Selection Fitness Trait Trait Negative Directional Selection Positive Directional Selection
Stabilizing selection Fitness average trait value
Malaria and Sickle Cell Anemia SSA caused by mutation in the past…
Disruptive Selection Fitness average trait value
Smith, T. B. 1990. Evolution. 44(4):832 • Black-bellied seed crackers (Pyrenestes) in Cameroon
skip Same birds different studyOpen portion of bar – Number hatchedBlack portion of bar – Number survived
skip Two more types of selection • Correlational – working on combinations of traits at once. (body size) (Wing length)
Frequency Dependent Selection – Level of selection dependent upon most common phenotype (at the time) Genyochromis Scale eater
Scalebiters – frequency dependent selection Right “mouthedness” Left “mouthedness”
Frequency of left “mouthedness” Frequency of right “mouthedness” Time
Selection can lead to speciation • Speciation: • Splitting of one species into two or more species (cladistic or punctuated). • Transformation of one species into a new species over time (gradual or anagenic).
B TIME A PHENOTYPIC VARIATION Anagenetic (gradual) versus Cladistic (punctuated) Speciation B A TIME A PHENOTYPIC VARIATION
Species • Many, many definitions • Biological species definition: A group of actually or potentially breeding individuals which are reproductively isolated from all other groups. • Doesn’t work in all situations • Fossils • Asexual organisms
Isolating mechanisms • Prezygotic – • Postzygotic -
Prezygotic I.M.s • Habitat isolation • Dendroica warblers in Eastern US
Modes of speciation • Allopatric – allo = different, patri = fatherland • Sympatric – sym = together • Has to do with location or habitat
Grand Canyon Kaibab Squirrel Abert’s Squirrel
Sympatric speciation • Rare • Polyploidy in plants • Hybrids
Divisions of Primates • Prosimians (“before ape”) • Lemurs • Tarsiers • Lorises • Galagos
Lemurs sifaka
Slow-loris galago
Old World Monkeys • Baboons • Macaques
Asian apes • Gibbons
African Apes • Chimps • Gorillas • Hominids • etc… bonobo