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Plant Organs: Leaves. Chapter 8. LEARNING OBJECTIVE 1. Describe the major tissues of the leaf ( epidermis, mesophyll, xylem, and phloem ) Relate the structure of the leaf to its function of photosynthesis. “Typical” Leaf. Blade. Veins. Petiole. Axillary bud. Stipules. Stem.
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Plant Organs: Leaves Chapter 8
LEARNING OBJECTIVE 1 • Describe the major tissues of the leaf (epidermis, mesophyll, xylem, and phloem) • Relate the structure of the leaf to its function of photosynthesis
Blade Veins Petiole Axillary bud Stipules Stem Fig. 8-1, p. 152
Animation: Simple and Compound Leaves CLICKTO PLAY
KEY TERMS • BLADE • Broad, flat part of a leaf • PETIOLE • Part of a leaf that attaches blade to stem
Simple Pinnately compound Palmately compound California white oak (Quercus lobata) Ohio buckeye (Aesculus glabra) White ash (Fraxinus americana) (a) Leaf form: simple and compound. Opposite Alternate Whorled American beech (Fagus grandifolia) Sugar maple (Acer saccharum) Southern catalpa (Catalpa bignonioides) (b) Leaf arrangement on a stem. Palmately netted Pinnately netted Parallel Sweetgum (Liquidambar styraciflua) Bermuda grass (Cynodon dactylon) Black willow (Salix nigra) (c) Venation patterns. Stepped Art Fig. 8-2, p. 154
KEY TERMS • PHOTOSYNTHESIS • The biological process that includes the capture of light energy and its transformation into chemical energy of organic molecules (such as glucose), which are manufactured from carbon dioxide and water
Animation: Leaf Organization CLICKTO PLAY
Epidermis • The transparent epidermisallows light to penetrate into the mesophyll, where photosynthesis occurs
KEY TERMS • CUTICLE • Waxy covering over epidermis of aerial parts (leaves and stems) of a plant • Enables the plant to survive in the dry conditions of a terrestrial environment
KEY TERMS • STOMA • Small pores in epidermis of stem or leaf • Permit gas exchange for photosynthesis and transpiration • Flanked by guard cells • GUARD CELL • Two guard cells form a pore (stoma)
Stomata • Stomata typically open during the day, when photosynthesis takes place, and close at night
KEY TERMS • MESOPHYLL • Photosynthetic ground tissue in the interior of a leaf • Contains air spaces for rapid diffusion of carbon dioxide and water into, and oxygen out of, mesophyll cells
Vascular Bundle • Leaf veinshave • xylemto conduct water and essential minerals to the leaf • phloemto conduct sugar produced by photosynthesis to rest of plant
KEY TERMS • BUNDLE SHEATH • One or more layers of nonvascular cells (parenchyma or sclerenchyma) surrounding the vascular bundle in a leaf
LEARNING OBJECTIVE 2 • Contrast leaf structure in eudicots and monocots
Animation: Monocot and Dicot Leaves CLICKTO PLAY
Upper epidermis Bundle sheath extension Bundle sheath Midvein Bundle sheath extension Lower epidermis Fig. 8-5, p. 157
Upper epidermis Palisade mesophyll Midvein Lengthwise view of vein Spongy mesophyll Privet Air space Lower epidermis Stoma Xylem Phloem (a) Privet (Ligustrum vulgare), a eudicot, has a mesophyll with distinct palisade and spongy sections. Fig. 8-6a, p. 158
Bundle sheath cells Midvein Mesophyll Parallel vein Upper epidermis Lower epidermis Phloem Xylem Fig. 8-6b, p. 158
Monocot and Eudicot Leaves • Monocot leaves • Usually narrow • Wrap around the stem in a sheath • Have parallel venation • Eudicot leaves • Usually have a broad, flattened blade • Have netted venation
Bulliform Cells • Large, thin-walled cellson upper epidermises of leaves of certain monocots (grasses) • Located on both sides of the midvein • May help leaf roll or fold inward during drought
(a) A folded leaf blade. The inconspicuous bulliform cells occur in the upper epidermis on both sides of the midvein. Bulliform cells Midvein Fig. 8-7a, p. 159
Bulliform cells (b) An expanded leaf blade. A higher magnification of the midvein region shows the enlarged, turgid bulliform cells. Mesophyll cell Midvein Fig. 8-7b, p. 159
LEARNING OBJECTIVE 3 • Outline the physiological changes that accompany stomatal opening and closing
Open Closed Guard cells Subsidiary cells (a) Guard cells of eudicots and many monocots are bean shaped. Fig. 8-8a, p. 160
Open Closed Guard cells Subsidiary cells (b) Some monocot guard cells (those of grasses, reeds, and sedges) are narrow in the center and thicker at each end. Fig. 8-8b, p. 160
Animation: Stomata CLICKTO PLAY
Stomatal Opening 1 1. Blue lightactivates proton pumps • in guard-cell plasma membrane • 2. Protons (H+) are pumped out of guard cells, forming a proton gradient • Charge and concentration difference on two sides of the guard-cell plasma membrane
KEY TERMS • PROTON GRADIENT • Difference in concentration of protons on the two sides of a cell membrane • Contains potential energy that can be used to form ATP or do work in the cell
Stomatal Opening 2 3. Gradient drives facilitated diffusionof potassium ions into guard cells 4. Chloride ions also enter guard cells through ion channels • Ions accumulate in vacuoles of guard cells • Solute concentration becomes greater than that of surrounding cells
KEY TERMS • FACILITATED DIFFUSION • Diffusion of materials from a region of higher concentration to a region of lower concentration through special passageways in the membrane
Stomatal Opening 3 • 5. Water enters guard cells from surrounding epidermal cells by osmosis • Increased turgidity changes the shape of guard cells, causing stoma to open
Blue light activates proton pumps. Protons are pumped out of guard cells, forming proton gradient. Potassium ions enter guard cells through voltage-activated ion channels. Chloride ions also enter guard cells through ion channels. Water enters guard cells by osmosis,and stoma opens. 1 2 3 4 5 Fig. 8-9, p. 162
Stomatal Closing • As evening approaches, sucrose concentration in guard cells declines • Sucrose is converted to starch (osmotically inactive) • Water leaves by osmosis, guard cells lose their turgidity, pore closes
Blade Petiole Fig. 8-10, p. 163
Guard cells of sunken stoma Epidermis and cuticle Resin duct Endodermis Xylem Vascular bundle Phloem Mesophyll cell (photosynthetic parenchyma cell) Fig. 8-11, p. 164
LEARNING OBJECTIVE 4 • Discuss transpiration and its effects on the plant