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Chapter 26. Phylogeny & The Tree of life. Phylogeny: Evolutionary history of a species or group of species Determined by evidences from fossil record , homologous structures, molecular homologies Systematics : Helps us understand phylogeny (data analysis of phylogeny).
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Chapter 26 Phylogeny & The Tree of life
Phylogeny: • Evolutionary history of a species or group of species • Determined by evidences from fossil record, homologous structures, molecular homologies • Systematics: • Helps us understand phylogeny (data analysis of phylogeny) Phylogeny & Systematics
Grouping according to evolutionary similarities • Binomial nomenclature (Linnaeus) • Domains Species • Taxon (plural: taxa): named taxonomic hierarchy • Ex: Panthera is the taxon at the genus level Taxonomy
Link phylogeny and taxonomy Phylogenic Trees
Phylogenies are inferred from morphological and molecular data 26.2
With ability to sequence DNA, we can easily show evolutionary relationships • EX: • Species 1: GAGATCTACACGGGGCCATGGAAAG • Species 2: GAGAACTACACGGGGCTATGGAAAG • Species 1: GAGATCTACACGGGGCCATGGAAAG • Species 3: GCCCACTATTATGGGCTATGGACCC DNA documentation
Shared characters are used to construct phylogenic trees 26.3
Science of constructing a cladogram Cladistics
Clades • Groups of organisms sharing a common ancestor Cladistics
A valid clade is monophyletic, it consists of the ancestor species and all its descendants Monophyletic
A paraphyletic clade consists of an ancestral species and some, but not all, of the descendants Paraphyletic
A polyphyletic clade includes many species that lack a common ancestor Polyphyletic
Characteristics • Shared primitive character • Homologous structure that is older than the branching of a particular clade from other members of that clade • It is shared by more than just the taxon we are trying to define. • Example – mammals all have a backbone, but so do other vertebrates.
Characteristics • Shared derived character • New evolutionary feature, unique to a particular group • Example - all mammals have hair, and no other animals have hair.
Ingroup & Outgroup In: Group of study (make comparisons) Out: Group that diverged prior to ingroup
An organism’s evolutionary history is documented in its genome 26.4
Orthologous genes • Same gene, found in different species • Divergence traces back to speciation event • Ex: cytochrome c • Paralogous genes • Same gene, found in same species Genome evolution: Gene duplication
“Time” evolution by looking at how genes change Example: HIV Molecular clocks
New information continues to revise our understanding of the tree of life 26.6