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Stems - I. Primary Structure of Stems. Dermal – Epidermis Ground Tissue Cortex Pith Other Vascular Tissue – vascular bundles Collateral VB Bicollateral VB Concentric VB. Acorus stem with endodermis. Amphivasal VB. Morphological concepts used to explain primary anatomy. Stelar Theory
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Primary Structure of Stems • Dermal – Epidermis • Ground Tissue • Cortex • Pith • Other • Vascular Tissue – vascular bundles • Collateral VB • Bicollateral VB • Concentric VB
Morphological concepts used to explain primary anatomy • Stelar Theory • Leaf Trace Theory
Types of steles Protosteles --------------> Siphonosteles-------------->
Protosteles have a central core of vascular tissue surrounded by cortex (not shown) Xylem Phloem
Siphonosteles have a central pith surrounded by a cylinder of vascular tissue Ectophloic Siphonostele Amphiphloic siphonostele Dictyostele
Ectophloic siphonstele in fern rhizome with a ring of xylem surrounded by a ring of phloem
Amphiphloic siphonostele in a fern rhizome with two rings of phloem on each side of xylem
Dictyosteles are considered amphiphloic siphonosteles with overlapping leaf gaps
Eustele Dicots and gymnosperms Monocots
Flattened protostele in Selaginella stem Flattened haplostele? OR Actinostele with two arms?
Actinostele with irregular shape Or a plectostele with connections between xylem areas?
Stele in stem Asteroxylon(fossil member of Division Lycophyta)
Anatomy of nodes Unilacunar with 2 traces Unilacunar with 1 trace Multilacunar with many gaps and many traces Trilacunar with 3 traces