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Chapter 50. An Introduction to Ecology and the Biosphere. Overview: The Scope of Ecology. Ecology is the scientific study of the interactions between organisms and the environment These interactions determine distribution of organisms and their abundance
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Chapter 50 An Introduction to Ecology and the Biosphere
Overview: The Scope of Ecology • Ecology is the scientific study of the interactions between organisms and the environment • These interactions determine distribution of organisms and their abundance • Ecology reveals the richness of the biosphere
Ecology and Evolutionary Biology • Events that occur in ecological time affect life on the scale of evolutionary time
Organisms and the Environment • The environment of any organism includes: • Abiotic, or nonliving, components • Biotic, or living, components • All the organisms that are part of the individual’s environment are collectively called the biota • Environmental components affect the distribution and abundance of organisms
LE 50-2 Kangaroos/km2 > 20 10–20 5–10 1–5 0.1–1 < 0.1 Limits of distribution Tasmania
Ecologists use observations and experiments to test explanations for the distribution and abundance of species • Organismal ecology studies how an organism’s structure, physiology, and (for animals) behavior meet environmental challenges
LE 50-3a Organismal ecology
Population ecology focuses on factors affecting how many individuals of a species live in an area Population ecology
Community ecology deals with the whole array of interacting species in a community Community ecology
Ecosystem ecology emphasizes energy flow and chemical cycling among the various biotic and abiotic components Ecosystem ecology
Landscape ecology deals with arrays of ecosystems and how they are arranged in a geographic region Landscape ecology
The biosphere is the global ecosystem, the sum of all the planet’s ecosystems • Ecology provides the scientific understanding underlying environmental issues • Rachel Carson is credited with starting the modern environmental movement
Most ecologists follow the precautionary principle regarding environmental issues • It states that humans need to be concerned with how their actions affect the environment
Concept 50.2: Interactions between organisms and the environment limit the distribution of species • Ecologists have long recognized global and regional patterns of distribution of organisms within the biosphere • Many naturalists began to identify broad patterns of distribution by naming biogeographic realms • Biogeography is a good starting point for understanding what limits geographic distribution of species
LE 50-5 Palearctic Nearctic Tropic of Cancer (23.5°N) Oriental Ethiopian Equator Neotropical (23.5°S) Tropic of Capricorn Australian
LE 50-6 Species absent because Area inaccessible or insufficient time Yes Yes Dispersal limits distribution? Habitat selection Yes Predation, parasitism, competition, disease Chemical factors Behavior limits distribution? No Biotic factors (other species) limit distribution? Water Oxygen Salinity pH Soil nutrients, etc. No Abiotic factors limit distribution? No Temperature Light Soil structure Fire Moisture, etc. Physical factors
Dispersal and Distribution • Dispersal is movement of individuals away from centers of high population density or from their area of origin • Dispersal contributes to global distribution of organisms • Natural range expansions show the influence of dispersal on distribution
LE 50-7 New areas occupied Year 1996 1989 1974
Species Transplants • Species transplants include organisms that are intentionally or accidentally relocated from their original distribution • Species transplants can disrupt the communities or ecosystems to which they have been introduced • Some organisms do not occupy all of their potential range • Species distribution may be limited by habitat selection behavior
Biotic Factors • Biotic factors that affect the distribution of organisms may include: • Interactions with other species • Predation • Competition
LE 50-8 100 Sea urchin Both limpets and urchins removed 80 Only urchins removed 60 Seaweed cover (%) Limpet 40 Only limpets removed Control (both urchins and limpets present) 20 0 August 1982 February 1984 August 1983 February 1983
Abiotic Factors • Abiotic factors affecting distribution of organisms: • Temperature • Water • Sunlight • Wind • Rocks and soil
Factors to Distribution • Environmental temperature is an important factor in distribution of organisms because of its effects on biological processes • Water availability in habitats is another important factor in species distribution • Light intensity and quality affects photosynthesis • Light is also important to development and behavior of organisms sensitive to photoperiod • Wind amplifies effects of temperature by increasing heat loss from evaporation and convection • Wind can change morphology of plants
Factors to Distribution • Many characteristics of soil (Rocks and Soil )limit distribution of plants and thus the animals that feed upon them: • Physical structure • pH • Mineral composition
Climate • Four major abiotic components of climate: temperature, water, sunlight, and wind • Climate is the prevailing weather in an area • Macroclimate consists of patterns on the global, regional, and local level • Microclimate consists of very fine patterns, such as those encountered by the community of organisms underneath a fallen log
Global Climate Patterns • Global climate patterns are determined largely by solar energy and the planet’s movement in space • Sunlight intensity plays a major part in determining the Earth’s climate patterns
LE 50-10a North Pole 60°N Low angle of incoming sunlight 30°N Tropic of Cancer Sunlight directly overhead at equinoxes 0° (equator) Tropic of Capricorn 30°S Low angle of incoming sunlight 60°S South Pole Atmosphere
Seasonal variations of light and temperature increase steadily toward the poles
LE 50-10b March equinox: Equator faces sun directly; neither pole tilts toward sun; all regions on Earth experience 12 hours of daylight and 12 hours of darkness. 60°N 30°N June solstice: Northern Hemisphere tilts toward sun; summer begins in Northern Hemisphere; winter begins in Southern Hemisphere. 0° (equator) 30°S December solstice: Northern Hemisphere tilts away from sun; winter begins in Northern Hemisphere; summer begins in Southern Hemisphere. Constant tilt of 23.5° September equinox: Equator faces sun directly; neither pole tilts toward sun; all regions on Earth experience 12 hours of daylight and 12 hours of darkness.
Global air circulation and wind patterns play major roles in determining climate patterns
LE 50-10c 60°N 30°N Descending dry air absorbs moisture Descending dry air absorbs moisture 0° (equator) Ascending moist air releases moisture 30°S 60°S 0° 23.5° 23.5° 30° 30° Arid zone Arid zone Tropics
LE 50-10d Arctic Circle 60°N Westerlies 30°N Northeast trades Doldrums 0° (equator) Southeast trades 30°S Westerlies 60°S Antarctic Circle
Regional, Local, and Seasonal Effects on Climate • Various features of the landscape contribute to local variations in climate • Seasonal variation also influences climate • Oceans and their currents and large lakes moderate the climate of nearby terrestrial environments
Mountains • Mountains have a significant effect on • The amount of sunlight reaching an area • Local temperature • Rainfall
LE 50-12 Wind direction East Pacific Ocean Sierra Nevada Coast Range
Seasonality • The angle of the sun leads to many seasonal changes in local environments • Lakes are sensitive to seasonal temperature change and experience seasonal turnover
LE 50-13 Spring Winter O2 (mg/L) O2 (mg/L) 0 4 8 12 0 4 8 12 Lake depth (m) Lake depth (m) 8 8 16 16 0° 4° 4° 2° 24 24 4° 4° 4° 4° 4° 4° 4°C 4°C O2 concentration High (>8 mg/L) Medium (4–8 mg/L) Low (<4 mg/L) O2 (mg/L) O2 (mg/L) 0 4 8 12 0 4 8 12 Lake depth (m) Lake depth (m) 4° 22° 8 8 4° 20° 18° 4° 8° 4° 16 16 6° 4° 5° 4°C 4°C 24 24 Summer Thermocline Autumn
Microclimate • Microclimate is determined by fine-scale differences in abiotic factors Long-Term Climate Change: • One way to predict future global climate change is to study previous changes
LE 50-14 Current range Predicted range Overlap 4.5°C warming over next century 6.5°C warming over next century
Concept 50.3: Abiotic and biotic factors influence the structure and dynamics of aquatic biomes • Varying combinations of biotic and abiotic factors determine the nature of biomes • Biomes are the major ecological associations that occupy broad geographic regions of land or water • Aquatic biomes account for the largest part of the biosphere in terms of area • They can contain fresh water or salt water • Oceans cover about 75% of Earth’s surface and have an enormous impact on the biosphere
LE 50-15 30°N Tropic of Cancer Equator Continental shelf Tropic of Capricorn 30°S Key Lakes Abyssal zone (below oceanic pelagic zone) Estuaries Rivers Oceanic pelagic zone Intertidal zone Coral reefs
Many aquatic biomes are stratified into zones or layers defined by light penetration, temperature, and depth
LE 50-16a Littoral zone Limnetic zone Photic zone Pelagic zone Benthic zone Aphotic zone Zonation in a lake
LE 50-16b Intertidal zone Neritic zone Oceanic zone 0 Photic zone 200 m Continental shelf Pelagic zone Benthic zone Aphotic zone 2,500–6,000 m Abyssal zone (deepest regions of ocean floor) Marine zonation
Major aquatic biomes: • Lakes • Wetlands • Streams and rivers • Estuaries • Intertidal zones • Oceanic pelagic biome • Coral reefs • Marine benthic zone