Phylum Echinodermata

1. Phylum Echinodermata

2. ECHINODERMS • Invertebrates • Inhabit marine environments ranging from coastal shallow waters to ocean trenches more than 10,000 m deep • Vary in diameter from 1cm to 1 m • Often brilliantly colored • Deuterostomes • unlike all other invertebrates • more closely related to chordates than other other invertebrates

3. HISTORY • More than 500 million year old • Early echinoderms appear to have been sessile & evolved form of radial symmetry as an adaptation to sessile existence • Later evolved ability to move from place to place • Vast species now moves by crawling SLOWLY along the ocean bottom • About 80 species are sessile • Approximately 6,000 total species of echinoderms have been classified

4. General Characteristics • Adults exhibit pentaradial symmetry – 5 points radiate from a central axis • Radial symmetry is secondary; larvae are bilaterally symmetrical and undergo metamorphosis to become radially symmetrical adults. Echinoderm larva

5. Poorly ganglionated; possess few sensory structures • Body wall contains an endoskeletonof calcareous plates - ossicles

6. General Characteristics cont. • Possess a network of canals throughout the body - water vascular system. • The canals are connected to extensions called tube feet (=podia),located on the oral surface • The water vascular system is important for locomotion, feeding, and gas exchange. http://www.youtube.com/watch?v=HPhAGyDceLo • Sexes are separate (dioecious), gametes (egg&sperm) shed into water, fertilization is external (SPAWNING)

7. Echinoderm CLASSES

8. Class Asteroidea • Sea stars (starfish) - means “starlike” • Typically have 5 arms which merge with a central disc • Mouth is located in the center of oral surface which is directed downward

9. Water Vascular System • On the aboral surface is the opening of the water vascular system the madreporite (=sieve plate) • Water enters the madreporite and goes through the stone canalcanal to the ring canal • Water then passes through a radial canalextending into each arm • All along the length of these canals are lateral canals that terminate in a bulb-like structures called ampullaeequipped with tube feet • Tube feet line the grooves on the oral surface - ambulacral grooves

10. Return to Taxonomy Asterias dissection 1 Oral surface Aboral surface

11. How the Podia Operate • Ampulla contract and force fluid into the podia causing it to become extended • Suckers at the tips of the podia come into contact with the substrate and adhere to the surface • Then the podia contract, thereby forcing water back into the ampulla, and the body is pulled forward

12. Nutrition • Mouth leads to a 2-part stomach: a large cardiac stomachand a smaller pyloric stomach • The pyloric stomach connects with digestive glands (=pyloric cecae)that runs into each arm • A short intestine extends from from the pyloric stomach to an anus on the aboral surface • Associated with the intestine are rectal cecae that pump the fecal wastes out of the anus

13. Return to Taxonomy Asterias dissection 3 • Aboral surface removed.

14. Additional Characteristics • The endoskelton is made up of calcareous plates that often penetrate the dermis as spines • Between the spines and plates are projections called papulae,which function in gas exchange and excretion • Other projections on the body wall include tiny jaw-like appendages called pedicellaria

15. Class Echinoidea • Sea urchins & sand dollars … means “spinelike” • Lack arms • Body is enclosed in a shell or test • Body surface is usually covered with moveable spines

17. Sea Urchins • Spherical body • Ambulacral plates bearing tube feet that radiate out toward the aboral surface • Use podia and spines during locomotion • The spines are moveable and articulate with the with the calcareous ossicles

18. Sea urchins generally feed by scraping algae off of rocks • Accomplished via a complex chewing apparatus called Aristotle's lantern

19. Class Holothuroidea • Means “water polyp” • Lack arms • Oral-aboral axis is greatly extended • Endoskeleton is reduced to a few ossicles scattered over the surface of the animal making them rather soft bodied • Some species crawl along the substrate using podia; others have peristaltic locomotion via muscle contractions Dermal ossicles

21. At the oral end of the body are a group of tentacles (modified podia) that surround the mouth; used in feeding • Have a muscular cloaca that is partly used in gas exchange • The actual gas exchange structures are branching structures called respiratory trees

22. ClassCrinoidea • Means “lily-like” • Most primitive of the echinoderms • Unusual in that the oral surface is directed upward • Aboral surface is attached to the substrate by means of a bendable stalk • The portion of the crinoid body attached to the stalk is called the crown; bears a number of arms

23. Along the length of the arms are branches called pinnules • The arms and the pinnules have ambulacral grooves with suckerless podia (secrete mucus) • The ambulacral grooves are heavily ciliated and the cilia is used to direct food to the mouth (=filter feeding)

24. Class Ophiuroidea“in the form of snake tail” • Includes basket stars and brittle stars • Has over 2,000 species • Most diverse group • Often overlooked because of size & tendency to occupy crevices • Arms are long and sharply set off from central disk • Madreporite is on the oral surface

25. Tube feet lack suction disks & locomotion occurs thru modification of skeleton that permits a unique form of grasping and movement