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Proximate and Ultimate Causes of Animal Behaviour. The beewolf wasp ( Philanthus triangulum ). Beewolf wasp burrows. Niko Tinbergen: one of the founders of ethology. The Scientific Method. Observation and causal question H ypothesis P redictions Test of predictions Conclusion.
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The Scientific Method • Observation and causal question • Hypothesis • Predictions • Test of predictions • Conclusion
The Scientific Method • Observation and causal question: Beewolves find the entrance to their own nests. How do they do it? • Hypothesis: They use local landmarks as cues • Predictions: If they use local landmarks then manipulating those landmarks should mislead them • Test of predictions: Experiments • Conclusion: Yes, local landmarks are used by beewolves as cues in finding own nest
Black-headed gull … and its chicks
The Scientific Method • Observationand causal question - black headed gulls remove eggshells. Why did black headed gulls evolve behaviour of eggshell removal? • Hypothesis - maybe this behaviour helps parent gulls to conceal their offspring from predators • Predictions - if so, broken eggshells should help predators find food • Test of predictions - experiment • Conclusion - yes, eggshell removal appears to help parent gulls conceal offspring from predators
Oystercatcher Kittiwake Black-headed gull
Oystercatcher Solitary nesting Nest predators Immediate eggshell removal Kittiwake Group nesting No nest predators No eggshell removal Black-headed gull Group nesting Nest predation present Delayed eggshell removal
Why do babies cry???