5 Basic types of species interactions

1. 1 XXXXXXXXXX ? 2 Competition is important...just not as important as predation 5 Basic types of species interactions

3. habitat food Competition ...common use of a limited resource limited What can be limiting....?

4. FUNDAMENTAL REALIZED food A B habitat 2 dimension, but there are more....

5. Between and within species?? Intraspecific competition Interspecific competition

6. Population Abundance Time How do we see it or measure it? • Individual Responses • Behavior – feeding rate • Physiology – growth rate • Morphological – body size • Population responses • Abundance • Distribution

7. 3 types of competition?? Interference competition Exploitation competition Apparent competition

8. Interference competition - occurs directly between individuals via aggression etc. when the individuals interfere with foraging, survival, reproduction of others, or by directly preventing their physical establishment in a portion of the habitat. (-) B A (-)

9. Exploitation competition - occurs indirectly through a common, limiting resource, which acts as an intermediate. Ex. depletes the amount of food or fill up all the available space. (-) B A (-) + + - - R

10. Apparent competition - occurs indirectly between two species which are both preyed upon by the same predator. B A B A - - - - + + + P P

11. http://video.google.com/videoplay?docid=-6431936272692514162

12. Resource partitioning Population Abundance Resource Gradient Competition's role in ecology and evolution "species packing" Population Abundance Resource Gradient

13. Lake Malawi Lake Tanganyika Cichlids exhibit remarkable evolutionary convergence. Similar ecomorphs have evolved repeatedly within different cichlid assemblages. All of the cichlids in the left-hand column are from Lake Tanganyika. All of the cichlids in the right-hand column are from Lake Malawi, and are more closely related to one another than to any species within Lake Tanganyika. Note the similarities among color patterns and trophic morphologies.

14. Utilize resources along a gradient (See Wooton) • Food particle size • Temperature • Habitat

15. Examples: Classic Werner and Hall Experiments Bluegill Green Sunfish Pumpkinseed

16. Examples: Ciscoes adaptive radiation (Lindsey 1981)

17. Population Abundance Time Modeling Competition • Lotka-volterra model • Competition coefficients • The equations • What the hell is an isocline, anyway? • 4 graphs. See Molles

18. Exotics species that affect the ecology of native Wisconsin Fishes Pelagic planktivores - Alewife, Rainbow smelt, white perch Benthic insectivores – round goby, common carp Pelagic piscivores – Pacific salmon, brown trout Pelagic/benthic parasite – Sea lamprey Stream insectivores – brown trout, rainbow trout Pelagic filter-feeders – silver carp, bighead carp

19. Exotics fish utilize different habitats And a variety of food sources

20. How do they get here? People Ships ballast water Canals Aquaculture

21. Aquaculture Silver carp Bighead carp Ballast water Alewife white perch Round goby Intentional Stocking Pacific Salmon Brown trout Rainbow trout Common carp Rainbow smelt Canals Sea lamprey Rainbow smelt

22. LAKE SUPERIOR

23. 3 aspects that make them so successful...? High fecundity – lots of eggs / individual Excellent competitors – better at gaining resources then native species Exploit a less well used resource

24. Detrimental to sport fisheries... round gobies to smallmouth bass nests http://people.cornell.edu/pages/gbs27/gbssmb.html

25. Detrimental to ....life as we know it on all lakes and rivers http://www.youtube.com/watch?v=2ChwJiKKBdA