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Speciation Ch. 21. How Do Species Form?. Microevolution promotes change in small population, when does enough change create a new separate species? Speciation process that forms new species from existing ones 2 forces isolate species: Geographic Lead to reproductive isolation Genetic
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How Do Species Form? • Microevolution promotes change in small population, when does enough change create a new separate species? • Speciation process that forms new species from existing ones • 2 forces isolate species: • Geographic • Lead to reproductive isolation • Genetic • Pairing of chromosomes limit mating, etc…
Categorizing Species • Linnaeus system Latin; name made genius and species nomenclature • Base of morphological species concept; connect species by physical traits • What is the major problem with the morphological species concept? • Polymorphic species might be confused as multiple species • More exact methods of classification must be used
Modern Classification • Biological species concept categorizes species on reproductive criteria • If members of two populations cannot breed or produce fertile offspring under normal natural conditions they are not the same species • If two population cannot exchange genetic information they are no longer connected • How do we categorize asexual species? • Genetic markers through DNA technology
Ancient Classification • How do we categorize exist species? • Fossil records combined with modern observations • Phylogenetic species concept using morphological and genetic sequence data fossil records are grouped with similar existing species
Geographic Speciation • Populations of a species spread across a wide range experience different environmental factors • The greater the factors, the great genetic and phenotypic variation can be seen • Subspecies variation in a species population that is easily categorized • Five subspecies of rat snake all have different color patterns • Subspecies can still interbreed when in contact
Ring Species • If two subspecies never come directly in contact and never directly interbreed, are they still the same species? • Yes! They can be connected through other subspecies • Ring species intermediate between two populations that allows for the mixing of their genes • California salamanders • Ring species may be showing the process of two separate populations finally dividing into two species
Reproductive Isolation Activity • Crystal and Donatello: • Explain ecological isolation • Serena and Bruce: • Explain temporal isolation • Nick and Papaya: • Explain behavioral isolation • Alex and Michael: • Explain mechanical isolation • Benjamin and Ezreal: • Explain gametic isolation • Fox and Lyn: • Explain all postzygotic mechanism • YOUR TURN!!! • I’m done talking. Your turn to teach each other. • Use the rest of the class and tonight to research your topic in pairs • Tomorrow we will form two groups (one member from each pair) and you will teach your topic to that group • Prepare examples, bring in visual material, and be ready to answer their questions • Here are the pairs and topics:
Types of Geographic Speciation • Allopatric Speciation • “different homeland” • Physical barrier separates populations • Two steps: 1) Physically separate; stops gene flow 2) Mutate over time to become reproductively isolated • Archipelagos are good examples; produce species clusters close related species from a recent common ancestor; Darwin’s Finches • Test speciation with second contact • Either groups are reproductively isolated or they produce hybrids in hybrids zones • Over time these zones may disappear; reinforcement
Types of Geographic Speciation 2) Parapatric Speciation • “beside homeland” • Speciation between adjacent populations; driven by hybrids having a lower fitness in the environment 3) Sympatric Speciation • “same homeland” • One population produces subgroups that separate with time • Insect species lives on one type of plant (host); a mutation in a part of the population causes a shift in the selected host • New population moves to new host (host race); now separated they start their path towards speciation • Polyploidy plants typically experience this type of speciation
Genes and Speciation • Genetic changes ultimately drive speciation; all other factors just help in the eventual changes in the populations gene pools • Genetic Divergence genetic differences that reproductively isolate populations • Swordtails and Platys hybrids produce lethal tumors; genetics wont let hybrids survive • Snail species mutation reverse the direction shells coil, can no longer physically breed • Sexual selection in birds changes in songs and dances make males of one population not attractive to females of another
Genes and Speciation 2) Polyploidy extra copies of chromosomes can reproductively isolate populations • Autopolyploidy full nondisjunction produces an unreduced gamete (diploid sex cell) • Grass, violets, chrysanthemums • Unreduced gametes mix to make tetraploid offspring • Can self-pollinate or mate with other tetraploid plants • Reproductively isolated from all non-polyploidy populations
Genes and Speciation 2) Polyploidy extra copies of chromosomes can reproductively isolate populations 2) Allopolyploidy two closely related species form a hybrid, which is normally sterile, BUT if self-fertilize after forming gametes through meiosis or gains the needed chromosomes, it can form a new species • Coffee, cotton, potatoes, tobacco • Very fast speciation
Genes and Speciation 3) Chromosome Alteration inversions, translocations, deletions, and duplications can all alter the genome enough to cause speciation • Humans (46) diverged from gorillas and chimps (48) possibly because we lost 2 chromosomes that fused into Chromosome #2 • Inversion at centromere on #2 also points to divergence between chimps/humans and gorillas/orangutans • Humans differ from chimps by 6 other inversions
Homework • Crystal and Donatello: • Explain conditions for making fossils/why so few fossils survive • Serena and Bruce: • Explain relative and absolute dating of fossils • Nick and Papaya: • Explain how we use fossils to map out life millions of years ago • Alex and Michael: • Explain the continental drift and how it effects speciation • Benjamin and Ezreal: • Explain slow gradual changes to the surface of the Earth • Fox and Lyn: • Explain sudden dramatic changes to the surface of the Earth • Suggested Homework: • Test Your Knowledge Ch 21 • Actual Homework (Wednesday): • Discuss the Concepts #3 • Interpret the Data • Apply Evolutionary Concepts • Homework for Tuesday’s class: • Summary of each type of reproductive isolation with examples • Be prepared to do the group sharing again with 22.1 and 22.2