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Photosynthesis Has Two Sets of Reactions. Light-dependent reactions takes place in the thylakoids ; cannot take place unless light is present, traps energy. Light-independent reactions take place in the stroma; can occur in either the light or the dark, fixes carbon.
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Photosynthesis Has Two Sets of Reactions • Light-dependent reactions takes place in the thylakoids; cannot take place unless light is present, traps energy. • Light-independent reactions take place in the stroma; can occur in either the light or the dark, fixes carbon.
A. Light-Independent Reactions • 1. The second stage of photosynthesis; light is not directly required. • 2. Require CO2, which enters through leaf and NADPH and ATP, which have been produced by light-dependent reactions.
1. Fixing Carbon Dioxide • a. CO2 fixation is the attachment of CO2 to an organic compound. • b. RuBP (ribulose bisphosphate) is a five-carbon molecule that combines with carbon dioxide.
2. Reducing PGA • a. Six-carbon molecule immediately breaks down, forms two PGA (3-phosphoglycerate[C3]) molecules.
2. Reducing PGA • b. Each of two PGA molecules undergoes reduction to PGAL in two steps. • c. Light-dependent reactions provide NADPH (electrons) and ATP (energy) to reduce PGA to PGAL.
3. Regenerating RuBP • a. Every three turns of Calvin cycle, five molecules of PGAL are used to re-form three molecules of RuBP (ribulose bisphosphate).
3. Regenerating RuBP • b. Every three turns of Calvin cycle, there is net gain of one PGAL molecule; five PGAL regenerate RuBP. • c.First molecule identified by Calvin was PGA [C3], a three-carbon product; Calvin cycle is also known as C3 cycle.
B. The Importance of PGAL • 1. PGAL (glyceraldehyde-3-phosphate) is product of Calvin cycle; is converted to many organic molecules.
Photosynthesis Takes Other Routes;C3 and C4 Plants • 1. In C3 plants, Calvin cycle fixes CO2 directly; first molecule following CO2 fixation is PGA, a C3 molecule. • 2. C4leaves fix CO2 by forming a C4 molecule prior to the involvement of the Calvin cycle.
C3 and C4 Plants • C3 Plants • Wheat • Rice • Oats • C4 Plants • Corn • Sugarcane • Bermuda grass
Photorespiration • a. In hot weather, stomates close to save water; CO2 concentration decreases in leaves; O2 increases. • b. In C3 plants, O2 competes with CO2, resulting in production of only one molecule of PGA. • c. Called "photorespiration" since oxygen is taken up and CO2 is produced; produces only one PGA.
C4 Plants Flourish When It Is Hot and Dry • Photorespiration does not occur in C4 leaves even when stomates are closed. • C4 plants have advantage over C3 plants: in hot and dry weather, photorespiration does not occur (e.g., bluegrass dominates lawns in early summer, crabgrass takes over in hot midsummer).
The End. Practice Quiz Chapter 10