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Two Stages of Photosynthesis. Light Reaction aka light dependent rxn. Calvin Cycle aka light independent reaction. Transforming light photons into ATP and the electron carrier NADPH. Use ATP/NADPH to make glucose. The Calvin Cycle : C 3 carbon fixation . In stroma of chloroplast.
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Two Stages of Photosynthesis Light Reaction aka light dependent rxn Calvin Cycle aka light independent reaction Transforming light photons into ATP and the electron carrier NADPH Use ATP/NADPH to make glucose
The Calvin Cycle : C3 carbon fixation In stroma of chloroplast Inputs CO2 ATP NADPH from air from light rxn e Output G3P: a 3C organic molecule used to make glucose, fructose, starch, cellulose…
The Calvin Cycle : C3 carbon fixation 3 turns of the cycle Rubisco enzyme RuBP ATP ATP Energy input Most G3P used to regenerate RuBP e- G3P one ‘extra’ G3P Energy stored
Calvin Cycle Accounting Per turn 3 turn 6 turns Consume CO2 1 3 6 Consume ATP 3 9 18 Consume NADPH 2 6 12 Produces G3P 1 2 Produces Glucose 1
http://www.science.smith.edu/departments/Biology/Bio231/calvin.htmlhttp://www.science.smith.edu/departments/Biology/Bio231/calvin.html http://faculty.nl.edu/jste/calvin_cycle.htm http://highered.mcgraw-hill.com/sites/0070960526/student_view0/chapter5/animation_quiz_1.html http://www.blinkx.com/watch-video/photosynthesis-bioflix-animation/zWVQ8TsGQEG7FGsd2MWPTw http://www.wiley.com/college/boyer/0470003790/animations/photosynthesis/photosynthesis.htm
This version of photosynthesis is used by 85% of plants - the C3 plantsC3 plants include grains, soy, legumes, cotton, most trees, and more Selection produces functional adaptations, but not necessarily at optimal efficiency What about the other 15%? i.e. the C4 and CAM plants
In hot, dry conditions, plant will undergo Photorespiration • to conserve water, leaf stomata close, preventing CO2 intake and O2 release. • as CO2 concentration in the chloroplasts drops, Rubisco fixes oxygeninstead of CO2. • Photorespiration wastes radiant energy no glucose is made, no ATP is made What’s a plant to do?
C4 Plantsseparate C-fixation and the Calvin cycle into different cells CO2 uptake in mesophyll cells Light reaction occurs O2 produced stays here 4C molecule – PEP - made PEP shuttles CO2 to bundle-sheath cells 4C 3C + CO2 used in Calvin cycle Therefore can keep stomata closed
C4 Plantscorn, sugarcane In mesophyll: H20 input light rxn O2 , PEP fixes C PEP yields CO2 In bundle-sheath cells: Calvin cycle G3P sugar
CAM Plants C-fixation and Calvin at different times of the day Avoiding evaporative loss • Stomates open at night • CO2 is converted to an acid and stored hottest, driest conditions • During the day, the acid is broken down • CO2 released to RUBISCO for photosynthesis CAM plants can ‘idle’ a while