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Outline. The Concept of the Community Diversity and Composition Models The Structure of Communities Island Biogeography Habitat and Ecological Niche Competition Between Populations Predator-Prey Interactions Symbiotic Relationships Community Development Community Biodiversity.
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Outline • The Concept of the Community • Diversity and Composition Models • The Structure of Communities • Island Biogeography • Habitat and Ecological Niche • Competition Between Populations • Predator-Prey Interactions • Symbiotic Relationships • Community Development • Community Biodiversity
Community Concept • An assemblage of populations interacting with one another within the same environment • Composition is a listing of various species in the community • Diversity includes both species richness and species diversity
Diversity and Composition Models • Gleason - Individualistic Model • Each population is there because its abiotic requirements are met • Clements - Interactive Model • Community is the highest level of organization • Dependent on biotic interactions
Island Biogeography • MacArthur and Wilson • Developed a general model of island biogeography • Explains and predicts how the community diversity of an island is affected by • Distance from the mainland, and • Size of the island
Community Structure • Competition • When two species compete, the abundance of both species is negatively impacted • Predation (or parasitism) • Expected to increase the abundance of the predator (or parasite) • And reduce the abundance of the prey (or host)
Habitat and Ecological Niche • Habitat • The area an organism lives and reproduces in • Ecological niche • The role a species plays in its community • Includes its habitat, and • Its interactions with other organisms • Fundamental niche - All conditions under which the organism can survive • Realized niche - Set of conditions under which it exists in nature
Competition Between Populations • Interspecific competition • Members of different species require the same resource • The supply of the resource is limited • Competitive Exclusion Principle • No two species can occupy the same niche at the same time • Resource Partitioning decreases competition • Can lead to character displacement
Niche Specialization AmongFive Species of Coexisting Warblers
Predator-Prey Interactions • Predation • One living organism, the predator, feeds on another, the prey • Predator is larger • Predator has lower reproductive rate • Prey usually entirely consumed • Presence of predators can decrease prey densities, and vice-versa
Predator-prey Interaction BetweenParamecium caudatum and Didinium nasutum
Prey Defenses • Prey defenses • Mechanisms that thwart the possibility of being eaten by a predator • Spines • Tough Epidermis • Poisonous Chemicals • Camouflage • Bright Coloration • Flocking Behavior
Mimicry • Mimicry • One species resembles another species • Mimicked species possesses an overt antipredator defense • Batesian Mimicry - Mimic lacks defense of the organism it resembles • Müllerian Mimicry - Mimic shares same protective defense
Symbiotic Relationships • Symbiosis • Interactions in which there is a close relationship between members of two species • Parasitism • Parasite derives nourishment from a host, and may use host as habitat and mode of transmission • Endoparasites • Ectoparasites
Commensalism • Symbiosis, cont. • Commensalism • A symbiotic relationship in which one species benefits and the other is indifferent • Remoras and Sharks • Many supposed examples may turn out to be mutualism or parasitism • Inferred amount of harm or benefit two species do to one another is subject to investigator bias
Mutualism • Symbiosis, cont. • Mutualism • A symbiotic relationship in which both members of the association benefit • Need not be equally beneficial to both species • Cleaning Symbiosis • Often help each other obtain food or avoid predation • Bacteria in human intestinal tract
Community Development • Ecological Succession • A predictable pattern of change in species replacements following a disturbance • Primary Succession occurs in areas where there is no soil formation • Secondary Succession begins in areas where soil is present • Pioneer Species
Succession Models • Facilitation Model • Each stage facilitates invasion and replacement by organisms of the next stage • Succession in a particular area will always lead to the same type of community • Climax Community
Succession Models • Inhibition Model • Colonists remain and inhibit growth of other plants until the colonists are damaged or die • Tolerance Model • Different types of plants can colonize an area at the same time • Chance determine which seeds arrive first
Community Diversity • Community stability can be recognized in three ways • Persistence through time • Resistance to change • Recovery once a disturbance has occurred
Intermediate Disturbance Hypothesis • If widespread disturbances occur frequently, diversity will be limited • If diversity is high, only moderate disturbances have been occurring with moderate frequency
Predation, Competition, and Biodiversity • Predation by a particular species may reduce competition and increases diversity • Such predators are referred to as keystone predators • Exotic species • May lead to unbridled competition • Resultant reduction in biodiversity
Review • The Concept of the Community • Diversity and Composition Models • The Structure of Communities • Island Biogeography • Habitat and Ecological Niche • Competition Between Populations • Predator-Prey Interactions • Symbiotic Relationships • Community Development • Community Biodiversity